Affinage

SNAP23

Synaptosomal-associated protein 23 · UniProt O00161

Length
211 aa
Mass
23.4 kDa
Annotated
2026-06-10
100 papers in source corpus 68 papers cited in narrative 68 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 9/9 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

SNAP-23 is a ubiquitously expressed, palmitoylated t-SNARE that serves as the non-neuronal counterpart of SNAP-25, forming the core of SNARE complexes that drive regulated and constitutive membrane fusion across many cell types (PMID:8663154, PMID:9407084). It binds multiple syntaxins (notably syntaxin-2, -4, -6, -11) and VAMPs in vitro and reconstitutes functional 20S SNARE complexes with α-SNAP and NSF (PMID:8663154, PMID:9693005, PMID:10051443, PMID:10036234, PMID:11001914); its C-terminal exon-encoded domain is specifically required for VAMP binding and for fusion competence, distinguishing it from the syntaxin-binding domain (PMID:10713150, PMID:11350976). Through these complexes SNAP-23 mediates exocytosis in mast cells, platelets, neutrophils, endothelial cells, alveolar type II cells, and secretory epithelia, and supports insulin-stimulated GLUT4 translocation in adipocytes, transferrin/recycling-vesicle fusion, MMP secretion driving invadopodial ECM degradation, and Fc-receptor phagocytosis (PMID:9727496, PMID:9891020, PMID:10648404, PMID:11001914, PMID:10329959, PMID:10713150, PMID:16118213, PMID:19910495, PMID:23087210, PMID:24807903, PMID:26266817, PMID:27697926). Membrane targeting proceeds through electrostatic anchoring by basic residues in its cysteine-rich region to acidic phosphoinositides followed by palmitoylation, which also confers ~3-fold greater lipid-raft enrichment than SNAP-25 owing to an extra raft-promoting cysteine (PMID:15542596, PMID:28240595). SNAP-23 activity is gated by phosphorylation: PKC/thrombin signaling phosphorylates Ser23/Thr24 to inhibit syntaxin-4 binding, while IKK2/IKKβ phosphorylation at Ser95/Ser120 promotes SNARE assembly and fusion and is required for mast-cell degranulation, platelet secretion, and inflammatory cytokine release (PMID:12930825, PMID:15611044, PMID:18692471, PMID:23613522, PMID:23960234). Phosphorylation also redirects SNAP-23 from the plasma membrane to granule or autophagosomal membranes, where phospho-SNAP-23 engages the STX17 complex to drive autophagosome–lysosome fusion (PMID:28784843, PMID:33213278). SNAP-23 is essential for life: germline deletion causes pre-implantation lethality and conditional loss blocks lymphocyte development and triggers fibroblast apoptosis (PMID:21479242, PMID:25706117). Beyond canonical fusion, SNAP-23 supports ATG9-dependent macroautophagy in adipocytes (its loss causing lipodystrophy via BAX activation), lipid droplet–mitochondria coupling, and postsynaptic NMDA-receptor surface delivery in dendritic spines (PMID:20118925, PMID:30102258, PMID:19524684). In pancreatic β-cells SNAP-23 acts paradoxically as an inhibitory SNARE, competing with SNAP-25 for Ca2+-channel access at fusion sites such that its depletion enhances insulin exocytosis (PMID:27697926, PMID:32051343).

Mechanistic history

Synthesis pass · year-by-year structured walk · 28 steps
  1. 1996 High

    Established that a ubiquitous SNAP-25 homolog exists to provide t-SNARE function outside neurons, answering whether non-neuronal cells use a dedicated fusion factor.

    Evidence Yeast two-hybrid and in vitro binding from human B lymphocytes showing 23-kDa protein binds multiple syntaxins and VAMPs

    PMID:8663154

    Open questions at the time
    • In vitro binding does not establish which syntaxin/VAMP pairings operate in vivo
    • No fusion or secretion phenotype tested at identification
  2. 1997 High

    Defined how SNAP-23 attaches to membranes and that this attachment is functionally coupled to its t-SNARE activity, plus that it resists BoNT/E unlike SNAP-25.

    Evidence Metabolic [3H]palmitate labeling with in vivo syntaxin-binding correlation; BoNT/E cleavage and rescue of insulin secretion in HIT cells

    PMID:10329400 PMID:9407084

    Open questions at the time
    • Exact palmitoylated cysteines and palmitoyltransferase not defined
    • Functional rescue relies on overexpression
  3. 1997 High

    Showed SNAP-23 is predominantly plasma-membrane localized and that its assembly with syntaxin-4 is negatively regulated by Munc18c, establishing a docking checkpoint.

    Evidence Subcellular fractionation/immunofluorescence in 3T3-L1 adipocytes; COS-cell co-expression and co-IP with Munc18c

    PMID:9020061 PMID:9168999

    Open questions at the time
    • Munc18c regulation shown in overexpression, not endogenous setting
    • Dynamics of redistribution upon stimulation not addressed here
  4. 1998 High

    Reconstituted the binary SNARE interactions and revealed phosphoregulation, showing PKA phosphorylation of syntaxin-4 disrupts SNAP-23 binding and that SNAP-23 lacks the VAMP-potentiating activity of SNAP-25.

    Evidence Quantitative in vitro binding with kinase treatment (PKA, CKII, PKC)

    PMID:9693005

    Open questions at the time
    • Performed in vitro; cellular relevance of syntaxin-4 phosphorylation not tested
    • Does not address SNAP-23's own phosphorylation
  5. 1998 High

    Demonstrated stimulus-dependent redistribution of SNAP-23 from plasma membrane to granules drives compound exocytosis, linking SNAP-23 trafficking to secretory mode.

    Evidence Permeabilized mast-cell exocytosis with immunofluorescence and functional antibody inhibition

    PMID:9727496

    Open questions at the time
    • Signal driving relocation not identified
    • Granule-resident partners during compound fusion not defined
  6. 1998 High

    Extended SNAP-23 function to constitutive membrane recycling and regulated water-channel trafficking, showing it operates beyond classic regulated secretion.

    Evidence SLO-permeabilized MDCK transferrin recycling with BoNT/E and antibody inhibition; immunoisolation of AQP2 vesicles

    PMID:9651373 PMID:9815132

    Open questions at the time
    • AQP2 colocalization lacks functional perturbation
    • Cognate VAMP/syntaxin for recycling fusion not pinned down
  7. 1999 High

    Positioned SNAP-23 in insulin-regulated GLUT4 exocytosis and reconstituted authentic 20S SNARE complexes with syntaxin-4 and VAMP2/3, NSF and α-SNAP.

    Evidence Microinjection of antibody/recombinant protein in adipocytes; 20S complex reconstitution and co-IP; botulinum toxin GLUT4 assays

    PMID:10051443 PMID:10329959 PMID:10497159 PMID:9153412

    Open questions at the time
    • Relative contribution of SNAP-23 vs cellubrevin in distinct fusion steps unresolved
    • Insulin did not alter complex stoichiometry, leaving the regulated step undefined
  8. 1999 High

    Identified syntaxin-11 as a SNAP-23 partner on late endosomes/TGN and uncovered phosphorylation- and cytoskeleton-based control of the cytosolic SNAP-23 reservoir.

    Evidence Yeast two-hybrid/co-IP for syntaxin-11; SNAK kinase phosphorylation of unassembled SNAP-23; vimentin co-IP with NEM perturbation

    PMID:10036234 PMID:10588641 PMID:11029050

    Open questions at the time
    • SNAK identity and physiological trigger not defined
    • Vimentin-reservoir model relies on detergent fractionation and pharmacology
  9. 2000 High

    Established SNAP-23 with syntaxin-2/-4 as the platelet secretory t-SNARE for alpha-, dense-core, and lysosomal granules, generalizing its exocytic role to blood cells and neutrophils.

    Evidence SLO-permeabilized platelet/neutrophil exocytosis with antibody/peptide inhibition, co-IP, toxin cleavage, granule-specific readouts

    PMID:10648404 PMID:10961877 PMID:11001914 PMID:9891020

    Open questions at the time
    • Selectivity for granule subtypes mechanistically unexplained
    • Stimulus-coupled regulation of complex assembly not addressed here
  10. 2001 High

    Mapped the C-terminal VAMP-binding domain as essential for fusion and extended SNAP-23 function to surfactant secretion, defining a domain-level requirement for secretion.

    Evidence Deletion mutagenesis with mast-cell exocytosis; antisense/peptide/antibody perturbation in alveolar type II cells

    PMID:10713150 PMID:11350976 PMID:15035620

    Open questions at the time
    • Structural basis of C-terminal VAMP recognition not resolved
    • Whether the domain requirement is universal across secretory systems not tested
  11. 2002 High

    Defined post-secretory and ultrastructural features: calpain cleavage of SNAP-23 after release and a predominantly plasma-membrane resting distribution by immuno-EM.

    Evidence Calpain cleavage assays with inhibitors in permeabilized platelets; immunonanogold EM and fractionation; kinesin coiled-coil binding

    PMID:12010801 PMID:12121992 PMID:12475239

    Open questions at the time
    • Functional consequence of calpain cleavage not directly demonstrated
    • Kinesin-SNAP-23 functional role not tested at this stage
  12. 2003 High

    Defined PKC phosphorylation of SNAP-23 at Ser23/Thr24 as an inhibitory input on syntaxin-4 binding and introduced regulatory partners snapin and tomosyn.

    Evidence 32P labeling, MS, PKC phosphorylation with phosphomimetic binding; snapin and b-tomosyn ternary-complex and GLUT4 assays

    PMID:12832401 PMID:12877659 PMID:12930825

    Open questions at the time
    • How inhibitory PKC sites are reconciled with secretion-promoting phosphorylation unresolved
    • Snapin and tomosyn cellular roles for SNAP-23 not fully tested
  13. 2004 High

    Identified IKK2/Ser95-Ser120 as the secretion-promoting phosphorylation, distinguished SNAP-23 from SNAP-25 by raft affinity and Ca2+/synaptotagmin selectivity, refining its specialized fusion properties.

    Evidence 32P/mutagenesis in mast cells and platelets; DRM fractionation with cys/phe mutagenesis; in vitro Ca2+-defined docking assay

    PMID:14742706 PMID:15542596 PMID:15611044

    Open questions at the time
    • Kinase identity at Ser95/Ser120 not yet assigned in 2004 studies
    • How raft enrichment couples to fusion not mechanistically linked
  14. 2005 Low

    Probed SNAP-23 interactions with Ca2+ channels and cholesterol-dependent t-SNARE clustering as caveolar fusion sites, linking lipid microdomain organization to fusion-site assembly.

    Evidence Yeast two-hybrid/co-IP with L-type Ca2+ channels; cell-free caveolar fusion with correlative fluorescence/immuno-EM

    PMID:15963257 PMID:16118213

    Open questions at the time
    • Ca2+-channel interaction lacks functional validation and rests on a single method pair
    • Functional role of t-SNARE clusters beyond colocalization not proven
  15. 2007 Medium

    Identified annexin A2 as a Ca2+-dependent SNAP-23 partner acting through the cysteine-rich region and documented a surface-exposed SNAP-23 pool, expanding regulatory and topological complexity.

    Evidence GST pulldown/co-IP with CRR mutagenesis and fusion-inhibition antibody; immuno-EM, flow cytometry, and enzymatic surface release

    PMID:17485553 PMID:17575076

    Open questions at the time
    • Functional significance of the extracellular SNAP-23 pool unestablished
    • Annexin A2 mechanism limited to alveolar type II cells
  16. 2009 High

    Broadened SNAP-23 roles to lipid-droplet/mitochondria coupling, MMP secretion driving invasion, and provided biophysical evidence that SNAP-23 SNARE complexes are less stable than SNAP-25 complexes.

    Evidence siRNA with beta-oxidation and in vitro LD-mitochondria assay; DN/RNAi/toxin with invasion and gelatin-degradation readouts; single-molecule force spectroscopy

    PMID:19273577 PMID:19524684 PMID:19910495

    Open questions at the time
    • Lower complex stability not directly tied to physiological fusion kinetics in cells
    • LD-mitochondria function shown in single fibroblast system
  17. 2010 High

    Revealed a postsynaptic role distinct from SNAP-25, showing SNAP-23 specifically controls NMDA-receptor surface expression and currents in dendritic spines.

    Evidence Conditional KO and shRNA with electrophysiology and surface-receptor assays in neurons

    PMID:20118925

    Open questions at the time
    • SNARE partners mediating NMDAR delivery not defined
    • Relationship to canonical exocytic role unclear
  18. 2011 High

    Established SNAP-23 as essential for life and cell survival, showing germline deletion causes pre-implantation lethality and conditional loss blocks lymphocyte development and triggers apoptosis.

    Evidence Germline Snap23-null embryo analysis; CD19-Cre and T-cell-Cre conditional KO; acute fibroblast depletion apoptosis assay

    PMID:21479242 PMID:25706117

    Open questions at the time
    • Molecular cause of essentiality (which fusion event is lethal to lose) not pinpointed
    • Apoptosis mechanism on acute depletion not defined here
  19. 2012 High

    Showed SNAP-23 supports phagosome maturation and ROS/acidification, with stimulus-dependent conformational opening monitored by intramolecular FRET requiring VAMP7.

    Evidence Overexpression/knockdown in macrophages, FRET probe, phagosome isolation, ROS/pH measurement

    PMID:23087210

    Open questions at the time
    • Trigger for conformational change beyond VAMP7 not defined
    • Which fusion events during maturation depend on SNAP-23 not fully dissected
  20. 2013 High

    Assigned IKKβ as the physiological kinase phosphorylating SNAP-23 at Ser95 to promote SNARE assembly and fusion, linking inflammatory signaling to secretion across platelets and mast cells.

    Evidence Platelet-specific IKKβ KO and inhibitors with proteoliposome fusion and SNARE co-IP; BMMC LPS/TNF assays with IKK-SNAP-23 co-IP

    PMID:23613522 PMID:23960234

    Open questions at the time
    • How phosphorylation mechanistically accelerates assembly at the molecular level not resolved
    • Coordination with inhibitory PKC sites unaddressed
  21. 2014 High

    Defined a SNAP23-syntaxin4-VAMP7 ternary complex driving MT1-MMP delivery to invadopodia and characterized synaptotagmin-7-triggered asynchronous release when SNAP-23 substitutes for SNAP-25.

    Evidence Co-IP/DN perturbation with invasion and degradation assays in breast cancer cells; viral rescue in SNAP-25/Syt7 double-KO autaptic neurons with electrophysiology

    PMID:24807903 PMID:25422940

    Open questions at the time
    • Why SNAP-23 yields asynchronous/spontaneous release at the molecular level not fully explained
    • Regulation of the invadopodial ternary complex beyond syntaxin-4 phosphorylation not detailed
  22. 2015 High

    Generalized SNAP-23 as the dominant secretory SNAP isoform in endothelial and airway secretory cells, mapped BoNT/A resistance determinants, and placed it downstream of the exocyst in recycling fusion.

    Evidence siRNA/KO with VWF and mucin secretion readouts; comprehensive mutagenesis/MD for BoNT/A resistance; siRNA transferrin recycling assay

    PMID:26092867 PMID:26182382 PMID:26266817 PMID:26523682

    Open questions at the time
    • Cell-type specificity of partner usage not unified into one model
    • Link between exocyst tethering and SNAP-23 SNARE assembly mechanistically thin
  23. 2016 High

    Revealed opposing tissue-specific functions in pancreas (promoting exocrine but restraining endocrine granule fusion) and identified a small molecule (MF286) that engages SNAP23 to promote insulin secretion.

    Evidence Pancreas-specific conditional KO with single-granule imaging; small-molecule screen/binding and in vivo insulin assay; septin7/NM-IIA complex co-IP and glucose uptake

    PMID:27697926 PMID:28011197

    Open questions at the time
    • Molecular basis of the exocrine-vs-endocrine reversal not resolved at this stage
    • MF286 mechanism of action on SNAP-23 not detailed
  24. 2017 High

    Defined the membrane-targeting sequence of events (electrostatic phosphoinositide anchoring before palmitoylation) and showed Rab5 and a basal/induced phosphorylation hierarchy control SNAP-23 membrane partitioning during exocytosis.

    Evidence Liposome reconstitution and basic-residue mutagenesis with live imaging; Rab5 siRNA exocytosis-mode assay; phospho-site mutant live imaging

    PMID:28240595 PMID:28784843 PMID:29127297

    Open questions at the time
    • Kinase for basal Thr102 site not identified
    • How Rab5 directs SNAP-23 to granules mechanistically unresolved
  25. 2018 High

    Connected SNAP-23 to ATG9-dependent macroautophagy (loss causing BAX-driven lipodystrophy) and to Kif5-mediated microtubule trafficking of mGlu1 receptors, broadening it from fusion executor to transport regulator.

    Evidence Adipocyte-specific KO with BAX/ATG9/ATG7 epistasis and lysosome inhibition; proteomics and recombination manipulation of SNAP23-Kif5 with trafficking imaging

    PMID:29762713 PMID:30102258

    Open questions at the time
    • SNARE partners for autophagic membrane fusion in adipocytes not fully mapped
    • Direct vs indirect role in microtubule transport not disentangled
  26. 2019 Medium

    Extended regulated exocytosis function to oocyte cortical granule release and meiotic arrest, and established syntaxin-11 cooperation in stimulus-dependent TLR4 surface delivery in macrophages.

    Evidence Trim-away depletion in oocytes with cortical granule/meiosis readouts; siRNA with TLR4 surface assay and intramolecular SNAP-23 FRET

    PMID:30811271 PMID:31201423

    Open questions at the time
    • Link between SNAP-23 secretion and gap-junction loss in oocytes mechanistically indirect
    • How syntaxin-11 drives SNAP-23 conformational change not resolved
  27. 2020 High

    Resolved the dual pancreatic role mechanistically: IKKβ-phosphorylated SNAP-23 relocates to autophagosomes to bind STX17 for autolysosome formation, while in β-cells SNAP-23 acts as an inhibitory SNARE competing with SNAP-25 for Ca2+ channels.

    Evidence shRNA in pancreatic slices with phospho-mutant analysis and STX17 co-IP plus pancreatitis model; β-cell KO/human islet knockdown with single-granule imaging and Ca2+-channel binding; NEDD4 ubiquitination co-IP

    PMID:32051343 PMID:33213278 PMID:33298171

    Open questions at the time
    • Stoichiometry of SNAP-23 vs SNAP-25 competition at Ca2+ channels not quantified
    • How phosphorylation switches SNAP-23 between STX4 and STX17 complexes not structurally defined
  28. 2022 Medium

    Defined further post-translational and signaling control: O-GlcNAcylation suppresses SNARE assembly/exosome release, and ICAM-1 clustering recruits SNAP-23 for localized chemokine secretion governing leukocyte transmigration route.

    Evidence OGT manipulation with SNARE co-IP and exosome/efflux assays; siRNA with chemokine release and transmigration assays

    PMID:34001861 PMID:35045291

    Open questions at the time
    • O-GlcNAc sites and their interplay with phosphorylation not mapped
    • Spatial recruitment mechanism downstream of ICAM-1 not resolved

Open questions

Synthesis pass · forward-looking unresolved questions
  • How the integrated post-translational code (palmitoylation, phosphorylation at multiple sites, O-GlcNAcylation, ubiquitination) and partner choice dictate when SNAP-23 acts as a fusion-promoting versus fusion-inhibiting SNARE, and how it switches between plasma-membrane, granule, and autophagosomal pools, remains unresolved.
  • No unified structural model linking modification state to complex partner selection
  • Quantitative competition with SNAP-25 across cell types not defined
  • Mechanism routing SNAP-23 between STX4 and STX17 complexes unknown

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0005198 structural molecule activity 5 GO:0008092 cytoskeletal protein binding 3 GO:0060090 molecular adaptor activity 3 GO:0008289 lipid binding 2
Localization
GO:0005886 plasma membrane 5 GO:0031410 cytoplasmic vesicle 4 GO:0005768 endosome 3 GO:0005811 lipid droplet 1 GO:0005856 cytoskeleton 1
Pathway
R-HSA-5653656 Vesicle-mediated transport 6 R-HSA-168256 Immune System 5 R-HSA-1430728 Metabolism 4 R-HSA-112316 Neuronal System 3 R-HSA-9609507 Protein localization 3 R-HSA-9612973 Autophagy 2
Complex memberships
20S SNARE complex (with NSF/alpha-SNAP)SNAP23-STX17 autophagosomal SNARE complexSNAP23-syntaxin4-VAMP2 SNARE complexSNAP23-syntaxin4-VAMP7 ternary complex

Evidence

Reading pass · 68 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
1996 SNAP-23 was identified as a 23-kDa protein from human B lymphocytes that binds to multiple syntaxins and synaptobrevins/VAMPs in vitro, establishing it as a ubiquitously expressed homolog of SNAP-25 and component of the t-SNARE complex in non-neuronal cells. Yeast two-hybrid screen, in vitro binding assays The Journal of biological chemistry High 8663154
1997 SNAP-23 is palmitoylated in vivo on cysteine residues in a central palmitoylation domain, and the extent of palmitoylation correlates with its ability to bind syntaxin in vivo. SNAP-23 is less palmitoylated than SNAP-25. Metabolic labeling with [3H]palmitate, in vivo binding studies Biochemical and biophysical research communications High 10329400
1997 SNAP-23 is resistant to cleavage by botulinum neurotoxin E (BoNT/E), unlike SNAP-25, and can functionally replace SNAP-25 to reconstitute insulin secretion in BoNT/E-treated HIT cells when overexpressed. BoNT/E cleavage assay, transfection rescue experiment in HIT cells The Journal of biological chemistry High 9407084
1997 Munc18c inhibits the binding of SNAP-23 to syntaxin 4 in a concentration-dependent manner, as shown in COS cell co-expression experiments. Co-expression in COS cells, co-immunoprecipitation Biochemical and biophysical research communications Medium 9168999
1997 SNAP-23 localizes predominantly to the plasma membrane in 3T3-L1 adipocytes and is membrane-bound across multiple non-neuronal tissues, with the majority associated with plasma membranes in subcellular fractionation. Subcellular fractionation, immunoblotting, immunofluorescence Biochemical and biophysical research communications High 9020061
1998 SNAP-23 forms binary, saturable interactions with syntaxin-4 and VAMP-2 in vitro. PKA phosphorylation of syntaxin-4 disrupts its binding to SNAP-23, whereas SNAP-23 does not potentiate VAMP-2 binding to syntaxin-4 (unlike neuronal SNAP-25). In vitro binding assays, phosphorylation by exogenous kinases (PKA, CKII, PKC) with binding readout Biochemistry High 9693005
1998 In response to stimulation, SNAP-23 relocates from plasma membrane lamellipodia-like projections to granule membranes in mast cells, and this relocation is required for compound exocytosis (granule-plasma membrane and granule-granule fusions). Permeabilized mast cell exocytosis assay, immunofluorescence, functional antibody inhibition Cell High 9727496
1998 SNAP-23 is required for transferrin recycling in MDCK cells: it is cleaved by BoNT/E in these cells, and addition of exogenous SNAP-23 or anti-SNAP-23 antibodies both inhibit ligand recycling in permeabilized cells. Streptolysin O-permeabilized MDCK cell reconstitution assay, BoNT/E cleavage, antibody inhibition The Journal of biological chemistry High 9651373
1998 SNAP-23 colocalizes with aquaporin-2 in collecting duct vesicles of rat kidney, and synaptobrevin-2 (VAMP-2) is also present in these AQP2-bearing vesicles, consistent with a role in vasopressin-regulated AQP2 trafficking. Subcellular fractionation, immunoisolation of AQP2-bearing vesicles, immunoblotting The American journal of physiology Medium 9815132
1999 SNAP-23 forms 20S SNARE complexes with syntaxin 4 and VAMP2/3 in rat adipose cell membranes with recombinant alpha-SNAP and NSF, and co-immunoprecipitates with syntaxin 4 in plasma membranes; insulin stimulation does not alter the SNAP-23/syntaxin 4 complex stoichiometry. Co-immunoprecipitation, 20S SNARE complex reconstitution with recombinant proteins The Biochemical journal High 10051443
1999 Syntaxin 11, identified by yeast two-hybrid screen using SNAP-23 as bait, binds SNAP-23 and VAMP in vitro and forms a complex with SNAP-23 in transfected HeLa cells and native B lymphocytes. Syntaxin 11 colocalizes with mannose 6-phosphate receptor on late endosomes and the trans-Golgi network. Yeast two-hybrid, in vitro binding, co-immunoprecipitation in transfected and native cells, immunofluorescence Journal of cell science High 10036234
1999 SNAP-23 is required for insulin-stimulated GLUT4 translocation to the plasma membrane: microinjection of anti-SNAP-23 C-terminal antibody inhibited GLUT4 membrane incorporation, while recombinant SNAP-23 enhanced the insulin effect. Microinjection of antibody and recombinant protein, permeabilized cell glucose transport assay The American journal of physiology High 10329959
1999 A novel kinase SNAK specifically phosphorylates SNAP-23 in vivo; only unassembled SNAP-23 (not in t-SNARE complexes) is phosphorylated, phosphorylated SNAP-23 resides in the cytosol, and SNAK co-expression enhances SNAP-23 stability and augments syntaxin assembly. In vivo phosphorylation assay, co-immunoprecipitation, co-expression studies Molecular biology of the cell High 10588641
1999 Botulinum toxin B (which cleaves cellubrevin/synaptobrevin) blocks insulin-stimulated GLUT4 translocation and glucose uptake in 3T3-L1 adipocytes; BoNT/A failed to cleave SNAP-23 and did not block glucose uptake, implicating cellubrevin rather than SNAP-23 proteolysis in one pathway of GLUT4 fusion. Toxin treatment of 3T3-L1 adipocytes, GLUT4 translocation assay, immunoblotting Biochemistry Medium 9153412
1999 Protein kinase B (PKB/Akt) stimulates GLUT4 translocation via a pathway involving SNAP-23 and synaptobrevin-2/cellubrevin; Clostridium botulinum toxin E (which cleaves SNAP-23) abolished PKB-DD-stimulated GLUT4 translocation but not insulin-stimulated GLUT1 or transferrin receptor translocation. Microinjection of botulinum toxins B and E into 3T3-L1 adipocytes, GLUT4 translocation assay The Journal of biological chemistry High 10497159
1999 SNAP-23 associates with vimentin intermediate filaments in a Triton X-100-insoluble fraction in fibroblasts. Upon NEM treatment, SNAP-23 dissociates from vimentin and forms a complex with syntaxin 4, suggesting vimentin-associated SNAP-23 serves as a reservoir for the plasma membrane fusion machinery. Detergent fractionation, co-immunoprecipitation, NEM treatment Molecular biology of the cell Medium 11029050
2000 SNAP-23 and syntaxin 4 are required for platelet alpha-granule release: they form a SDS-resistant heterotrimeric complex with VAMP that disassembles upon platelet activation, and antibodies to syntaxin 4 inhibited ~75% of Ca2+-induced alpha-granule release. Streptolysin O-permeabilized platelet exocytosis assay, co-immunoprecipitation, antibody inhibition, tetanus toxin cleavage The Journal of biological chemistry High 9891020
2000 SNAP-23 antibodies and C-terminal SNAP-23 peptide inhibit dense core granule release from platelets; SNAP-23 forms a complex in vivo with syntaxin 2, and anti-syntaxin 2 antibody also blocked dense core granule release. Streptolysin O-permeabilized platelet exocytosis assay, antibody/peptide inhibition, co-immunoprecipitation Blood High 10648404
2000 SNAP-23 mediates specific (CD67-enriched) granule secretion in human neutrophils but has little effect on azurophilic (CD63-enriched) granule exocytosis; SNAP-23 translocates to the cell surface upon activation and interacts with syntaxin 6 in vitro and in vivo, with interaction dramatically increased upon activation. Electropermeabilized neutrophil exocytosis assay, antibody inhibition, in vitro binding, co-immunoprecipitation Blood High 11001914
2000 SNAP-23 wild-type promotes the interaction between syntaxin 4 and VAMP2 both in vitro and in vivo; a C-terminal deletion mutant SNAP-23-ΔC8 that binds syntaxin 4 but not VAMP2 failed to mediate this interaction and inhibited insulin-induced GLUT4 but not GLUT1 translocation when overexpressed in adipocytes. In vitro binding, co-immunoprecipitation, adenovirus-mediated overexpression in 3T3-L1 adipocytes, GLUT4 translocation assay The Journal of biological chemistry High 10713150
2000 Platelet lysosome exocytosis requires both syntaxin 2 and syntaxin 4 together with SNAP-23 as t-SNARE components, in addition to NSF and ATP; these localize to granule membranes and open canalicular system respectively. Streptolysin O-permeabilized platelet exocytosis assay, antibody inhibition, localization studies Blood High 10961877
2001 The C-terminal domain of SNAP-23 (encoded by the last exon, 23 amino acids) is required for binding to VAMP but not syntaxin; overexpression of the carboxyl-terminal VAMP-binding mutant failed to enhance mast cell exocytosis, while wild-type SNAP-23 significantly enhanced it. Deletion mutagenesis, in vitro binding assays, overexpression in RBL mast cells with exocytosis assay The Journal of biological chemistry High 11350976
2001 SNAP-23 is required for regulated surfactant secretion in alveolar type II cells; SNAP-23 and syntaxin 2 both associate with plasma membrane and lamellar bodies, and peptides or antibodies targeting either protein inhibit Ca2+- and GTPγS-stimulated surfactant secretion from permeabilized cells. Antisense oligonucleotide knockdown, peptide/antibody inhibition in permeabilized type II cells, subcellular fractionation Biochemistry High 15035620
2001 SNAP-23 mediates H+-ATPase exocytosis to the apical membrane in inner medullary collecting duct (IMCD) cells; botulinum toxin A or E cleaves SNAP-23, reduces 20S SNARE complex formation, impairs H+-ATPase apical translocation by ~52%, and reduces H+ secretion by ~77%. Botulinum toxin cleavage, SNARE complex immunoprecipitation, H+-ATPase translocation and H+ secretion assays American journal of physiology. Cell physiology High 11245593
2002 Calpain specifically cleaves SNAP-23 (but not syntaxins 2 and 4 or alpha-SNAP) in activated platelets in a Ca2+-dependent and time-dependent manner; cleavage site is in the C-terminal third of the molecule; this occurs after granule release and may affect granule membrane exteriorization. In vitro calpain cleavage assay in permeabilized platelets, calpain inhibitors (calpeptin, calpastatin), Ca2+ dose-response The Journal of biological chemistry High 12121992
2002 SNAP-23 localizes predominantly to the plasma membrane in resting platelets (by immunonanogold electron microscopy), whereas human cellubrevin localizes ~80% to granule membranes and syntaxin 2 is distributed among alpha-granules, OCS, and plasma membrane. Pre-embedding immunonanogold electron microscopy, immunoblotting of subcellular fractions Blood High 12010801
2002 Kinesin heavy chain (residues 814–907) binds to SNAP-23 and SNAP-25 via coiled-coil interactions; the minimal SNAP23-binding region on kinesin is its cargo-binding domain, and a complex of kinesin heavy and light chains retains the ability to interact with SNAP-23. Yeast two-hybrid, in vitro and in vivo binding assays Biochemistry Medium 12475239
2003 SNAP-23 is phosphorylated in thrombin-activated platelets predominantly on serine residues via a PKC mechanism; phosphorylation at Ser23/Thr24 inhibits syntaxin 4 interactions; SNAP-23 phosphorylation kinetics parallel or precede granule secretion. 32P metabolic labeling, tryptic peptide mapping, mass spectrometry, PKC phosphorylation of recombinant SNAP-23, phosphomimetic mutant binding assays The Journal of biological chemistry High 12930825
2003 Snapin interacts with SNAP-23 (the ubiquitous SNAP-25 homolog) via its C-terminal helical domain and forms a ternary complex with SNAP-23 and syntaxin 4 in non-neuronal cells. Protein-protein interaction assays, deletion mapping, subcellular localization with GFP fusion, co-immunoprecipitation The Biochemical journal Medium 12877659
2003 b-Tomosyn forms a high affinity ternary complex with Syntaxin4 and SNAP23 that is competitively inhibited by VAMP-2; overexpression of Tomosyn in 3T3-L1 adipocytes inhibited GFP-GLUT4 translocation to the plasma membrane. In vitro binding assay, yeast two-hybrid, 3T3-L1 adipocyte overexpression with GLUT4-GFP translocation assay The Journal of biological chemistry Medium 12832401
2004 SNAP-23 phosphorylation at Ser95 and Ser120 is induced during mast cell degranulation and mouse platelet activation; overexpression of phosphorylation-deficient SNAP-23 mutants inhibits exocytosis; essentially all SNAP-23 in SNARE complexes (with syntaxin 4 and VAMP-2) is phosphorylated after stimulation. 32P metabolic labeling, site-directed mutagenesis, overexpression in RBL mast cells, co-immunoprecipitation The Journal of biological chemistry High 15611044
2004 SNAP-23 and SNAP-25 display different affinities for lipid raft microdomains: SNAP-23 (~54% raft-associated) is ~3-fold more enriched in rafts than SNAP-25 (~20%), due to substitution of a conserved phenylalanine in SNAP-25 with a cysteine in SNAP-23 that provides an additional palmitoylation site enhancing raft association. Detergent-resistant membrane isolation, site-directed mutagenesis of cysteine/phenylalanine residues The Journal of biological chemistry High 15542596
2004 SNAP-23 mediates granule docking at low Ca2+ (100 nM), whereas SNAP-25 mediates docking at high Ca2+ (1 μM), by interacting with different synaptotagmin family members; in intact endocrine cells, exogenous SNAP-23 increases basal (unstimulated) hormone secretion. In vitro granule docking assay with defined Ca2+ levels, overexpression in intact endocrine cells Molecular biology of the cell Medium 14742706
2005 SNAP-23 directly interacts with L-type Ca2+ channels in pancreatic acinar cells, demonstrated by yeast two-hybrid and immunoanalysis. Yeast two-hybrid, co-immunoprecipitation/immunoanalysis Journal of cellular and molecular medicine Low 15963257
2005 SNAP-23 and syntaxin-4 form cholesterol-dependent clusters on the endothelial basolateral plasma membrane, and ~50% of caveolae co-localize with these t-SNARE clusters during fusion, indicating these clusters are fusion sites for caveolar exocytosis. Cell-free caveolar fusion assay with plasma membrane sheets, correlative fluorescence microscopy and immunonanogold electron microscopy The Journal of biological chemistry High 16118213
2007 Annexin A2 binds SNAP-23 in a Ca2+-dependent manner via the cysteine-rich region (CRR) of SNAP-23; mutations of cysteines in the CRR dramatically reduced binding; anti-SNAP-23 antibody inhibited annexin A2-mediated fusion of lamellar bodies with the plasma membrane in alveolar type II cells. GST pulldown, co-immunoprecipitation, deletion/mutation analysis, in vitro fusion assay with inhibitory antibody American journal of respiratory cell and molecular biology High 17575076
2007 SNAP-23 and syntaxin-2 are expressed on the extracellular surface of the platelet plasma membrane; acyl-protein thioesterase 1 and botulinum toxin C light chain release SNAP-23 and syntaxin-2 respectively from intact platelet surfaces, and extracellular SNAP-23 retains the ability to bind syntaxin-2. Immunonanogold electron microscopy, flow cytometry of intact platelets, selective enzymatic release (APT1, BoNT/C), co-immunoprecipitation of released proteins Blood Medium 17485553
2008 IKK2 (IκB kinase 2) phosphorylates SNAP-23 in mast cells upon FcεRI stimulation; IKK2-deficient mast cells show impaired IgE-mediated degranulation, and ectopic expression of a phospho-mimetic SNAP-23 mutant partially rescues this defect. This function is NF-κB-independent. IKK2 knockout mice, in vitro degranulation assay, phospho-mimetic mutant rescue, in vivo anaphylaxis model Cell High 18692471
2009 SNAP-23 mediates lipid droplet–mitochondria complex formation in NIH 3T3 fibroblasts; siRNA ablation of SNAP23 reduced LD-mitochondria complex formation and beta-oxidation in an in vitro system of purified LD and mitochondria. siRNA knockdown, confocal and electron microscopy, in vitro LD-mitochondria interaction assay, beta-oxidation measurement Cell biology international Medium 19524684
2009 SNAP-23 in the SNARE complex containing SNAP-23A has less than half the stability of the complex containing SNAP-25B, as measured by single-molecule force spectroscopy; both create a local interaction at the ionic layer cuffing syntaxin 1A and synaptobrevin 2, positioning vesicles at ~13 nm from the plasma membrane. Single molecule force spectroscopy (AFM), ternary SNARE complex reconstitution The Journal of physiology Medium 19273577
2009 VAMP3, SNAP23, and syntaxin-13 colocalize with MMP2 and MMP9; dominant-negative SNARE mutants, RNAi, or tetanus toxin targeting these SNAREs impaired MMP secretion, MT1-MMP cell surface delivery, gelatin substrate degradation, and cell invasion. Dominant-negative SNARE mutants, RNAi, tetanus toxin, gelatin degradation assay, invasion assay Journal of cell science High 19910495
2009 SNAP-23 has punctate perinuclear localization overlapping with endosomal/transferrin receptor compartments in HepG2 and HT4 cells, distinct from the predominant plasma membrane pool, and redistributes to the apical domain as HepG2 cells polarize. Immunofluorescence, GFP-SNAP-23 expression, subcellular localization in polarizing cells Biochemical and biophysical research communications Medium 10049710
2010 SNAP-23 is enriched in dendritic spines and colocalizes with postsynaptic density components in neurons; loss of SNAP-23 (by KO mice or shRNA) markedly decreases NMDA receptor surface expression and NMDA receptor currents, whereas SNAP-25 loss does not affect postsynaptic NMDA receptors. Conditional knockout mice, shRNA knockdown, immunofluorescence, electrophysiology, surface receptor assay Nature neuroscience High 20118925
2011 SNAP-23 deletion results in pre-implantation embryonic lethality in mice; Snap23-null blastocysts die prior to implantation at E3.5, demonstrating SNAP-23 is essential for early embryogenesis. Homologous recombination to generate Snap23-null mice, analysis of pre-implantation embryos PloS one High 21479242
2011 Conditional deletion of SNAP-23 in CD19-Cre mice prevents B lymphocyte development; deletion using T lymphocyte-specific Cre mice prevents T lymphocyte development; acute depletion of SNAP-23 in fibroblasts leads to rapid apoptotic cell death. Conditional Cre-lox knockout mice, lymphocyte development assays, fibroblast apoptosis assay PloS one High 25706117
2012 SNAP-23 overexpression enhances Fc receptor-mediated phagocytosis in macrophages and promotes ROS production and acidification in phagosomes; SNAP-23 knockdown delays phagosome maturation and reduces uptake. SNAP-23 undergoes conformational change (increased FRET) on phagosomes specifically when VAMP7 is coexpressed. Stable cell line overexpression, siRNA knockdown, FRET probe, phagosome isolation, ROS and pH measurements Molecular biology of the cell High 23087210
2013 IKKβ phosphorylation of SNAP-23 at Ser95 controls platelet secretion; IKKβ inhibition or platelet-specific IKKβ knockout blocked SNAP-23 phosphorylation, platelet secretion, and SNARE complex formation; SNAP-23 phosphorylation enhanced membrane fusion of SNARE-containing proteoliposomes. IKKβ inhibitors, platelet-specific IKKβ-flox/flox × PF4-Cre mice, proteoliposome fusion assay, SNARE complex co-IP Blood High 23613522
2013 IKK-dependent phosphorylation of SNAP-23 is required for LPS-induced TNF secretion in mast cells; morphine prevents this by blocking IKK activation via a β-arrestin-2/TRAF6 complex, and SNAP-23 forms a complex with IKK in LPS-activated BMMCs. Bone marrow-derived mast cell activation assay, phosphorylation assays, co-immunoprecipitation, siRNA knockdown, pharmacological inhibition Journal of immunology Medium 23960234
2014 During invadopodium formation in MDA-MB-231 breast cancer cells, SNAP23, Syntaxin4, and VAMP7 form a ternary complex (detected by co-IP) that is required for MT1-MMP trafficking to invadopodia, ECM degradation, and cell invasion; increased SNAP23-Stx4-VAMP7 interaction correlates with decreased Syntaxin4 phosphorylation. Co-immunoprecipitation, dominant-negative SNARE perturbation, MT1-MMP localization assay, gelatin degradation, invasion assay Molecular biology of the cell High 24807903
2014 When SNAP-23 replaces SNAP-25 in hippocampal neurons, release becomes asynchronous and is triggered by endogenous synaptotagmin-7 (a plasma membrane Ca2+ sensor); SNAP-23-driven release is more asynchronous and has higher spontaneous release rates than SNAP-25-driven release. Viral transduction into SNAP-25/synaptotagmin-7 double-KO mouse autaptic neurons, electrophysiology, pHluorin imaging PloS one High 25422940
2015 SNAP23 is the predominant SNAP isoform in endothelial cells and localizes to the plasma membrane; SNAP23 knockdown decreased endothelial exocytosis of von Willebrand Factor and SNAP23 forms complexes with other known endothelial SNARE molecules. siRNA knockdown, VWF exocytosis assay, co-immunoprecipitation, immunofluorescence localization PloS one Medium 26266817
2015 SNAP23 is selectively expressed in airway secretory (goblet) cells and is required for stimulated mucin secretion; heterozygous SNAP23 knockout mice show impaired acute stimulated mucin release from perfused tracheas. Immunohistochemistry/FACS for cell-type selectivity, heterozygous Snap23 KO mouse trachea perfusion assay, static imaging assay Bioscience reports Medium 26182382
2015 BoNT/A resistance of SNAP-23 is conferred by 10 specific residues differing from SNAP-25; replacing these with SNAP-25 counterparts renders SNAP-23 cleavable by BoNT/A; conversely, transferring any of these SNAP-23 residues to SNAP-25 drastically decreases BoNT/A cleavability. Site-directed mutagenesis, BoNT/A cleavage assay, yeast-based screening, molecular dynamics simulation Journal of molecular biology High 26523682
2015 SNAP23 and VAMP2 mediate the fusion of transferrin receptor-containing recycling vesicles with the plasma membrane downstream of the exocyst complex; depletion of SNAP23 and/or SNAP25 suppresses recycling vesicle fusion, causing accumulation at the cell periphery. siRNA depletion, transferrin recycling assay, immunofluorescence colocalization Biology open Medium 26092867
2016 Loss of SNAP-23 in exocrine pancreas decreases stimulated granule fusion, while loss in endocrine pancreas increases granule fusion; MF286, a small molecule identified to bind specifically to SNAP23, promotes insulin secretion in mice. Pancreas-specific SNAP23 conditional KO mice, single granule imaging, small molecule screen and binding assay, in vivo insulin secretion assay The Journal of cell biology High 27697926
2016 Septin 7 forms a complex with NM-IIA and SNAP23; insulin decreases septin 7 and increases NM-IIA activity (myosin regulatory light chain phosphorylation) in the SNAP23 complex; septin 7 knockdown increases NM-IIA activity in the complex; NM-IIA activity in the SNAP23 complex promotes GSV docking/fusion with the plasma membrane in podocytes. Co-immunoprecipitation, siRNA knockdown, myosin light chain phosphorylation assay, glucose uptake assay Experimental cell research Medium 28011197
2017 Electrostatic anchoring by basic amino acids in the cysteine-rich region of SNAP-23 (and SNAP-25) mediates initial plasma membrane targeting through interaction with acidic lipids (phosphoinositides), prior to palmitoylation and stable membrane attachment. Site-directed mutagenesis of basic residues, liposome binding reconstitution assay, live cell imaging in neuroendocrine cells eLife High 28240595
2017 Rab5 is required for SNAP23 association with secretory granules in stimulated mast cells, and is pivotal for SNAP23-mediated homotypic granule-granule fusion (compound exocytosis); Rab5 silencing shifts secretion from compound to full (single granule) exocytosis mode. Rab5 siRNA silencing, live cell imaging, immunofluorescence, IKKβ2 inhibition Scientific reports Medium 29127297
2017 Phosphorylation at Thr102 of SNAP-23 (basal site) is required for initial plasma membrane association; induced phosphorylation at Ser95 and Ser120 promotes SNAP-23 internal membrane (granule membrane) association during mast cell exocytosis. GFP-SNAP-23 phosphorylation site mutant overexpression in RBL mast cells, live fluorescence imaging Biology open Medium 28784843
2018 Adipocyte-specific SNAP23 KO causes lipodystrophy through inhibition of ATG9-dependent (but ATG7-independent) macroautophagy, leading to failure of lysosomal BAX degradation, BAX activation, and apoptosis; BAX deficiency suppresses the lipodystrophic phenotype in SNAP23-KO mice. Adipocyte-specific conditional KO mice, BAX/ATG9/ATG7 co-deletion epistasis, lysosome inhibitor treatment, BAX colocalization with autophagic vacuoles The Journal of clinical investigation High 30102258
2018 SNAP23 forms a complex with Kif5 kinesin and directly interacts with the carboxyl terminus of the mGlu1 receptor; disrupting or enhancing the SNAP23-Kif5 interaction alters mGlu1 receptor trafficking along microtubules and its cell surface expression in hippocampal neurons. Proteomics (co-IP/MS), direct interaction assay, recombination approach to alter SNAP23-Kif5 interaction, FRAC (fluorescence recovery after cleavage) imaging Journal of molecular cell biology Medium 29762713
2019 SNAP23 is required for cortical granule exocytosis (regulated exocytosis) and for meiotic arrest in mouse oocytes; SNAP23 depletion (by Trim-away) causes premature meiotic resumption associated with loss of gap junction communication between oocyte and follicle cells. Trim-away protein depletion in oocytes, lectin staining/ZP2 cleavage for cortical granule exocytosis, meiotic arrest assay Biology of reproduction Medium 31201423
2019 Syntaxin 11 cooperates with SNAP-23 to regulate stimulus-dependent transport of TLR4 to the plasma membrane in macrophages; stx11 or SNAP-23 knockdown impairs TLR4 replenishment after LPS stimulation; stx11 knockdown reduces the FRET efficiency of an intramolecular SNAP-23 FRET probe induced by LPS stimulation. siRNA knockdown, TLR4 surface expression assay, intramolecular FRET probe for SNAP-23 conformational state Molecular biology of the cell Medium 30811271
2020 Pancreatic SNAP23 phosphorylation at Ser95/Ser120 (rat) or Ser120 (human) by IKKβ causes SNAP23 translocation from plasma membrane to autophagosomes where it binds the STX17 SNARE complex to regulate autophagosome-lysosome fusion; phosphorylation-disabled SNAP23 (S95A/S120A) cannot bind STX17; pancreatic SNAP23 knockdown impairs both STX4-driven basolateral exocytosis and STX17-driven autolysosome formation, protecting rats from pancreatitis. adenovirus-shRNA knockdown in rodent and human pancreatic slices and in vivo, phospho-mutant analysis, SNARE complex co-IP, in vitro pancreatitis model Autophagy High 33213278
2020 SNAP23 depletion in β cells paradoxically increases biphasic insulin secretion and exocytosis of both predocked and newcomer granules; in the context of β cells, SNAP23 acts as an inhibitory/weak partial fusion agonist by competing with SNAP25 for binding to Ca2+ channels at granule fusion sites. β-cell-specific conditional KO in mice, human islet viral knockdown (including T2D patients), single granule imaging, Ca2+ channel binding assay JCI insight High 32051343
2020 NEDD4 (an E3 ubiquitin ligase) binds SNAP23 and promotes its ubiquitin-mediated proteasomal degradation; BTX-A reduces SNAP23 expression by inhibiting IKKα/β phosphorylation and enhancing NEDD4-mediated ubiquitination of SNAP23. Co-immunoprecipitation of NEDD4-SNAP23, proteasome inhibitor rescue, IKKα/β phosphorylation assay Cell & bioscience Medium 33298171
2021 Reduced O-GlcNAcylation of SNAP-23 (caused by OGT downregulation) promotes formation of the SNARE complex (SNAP-23/VAMP8/Stx4), enhancing exosome release and cisplatin efflux in ovarian cancer cells; O-GlcNAcylation of SNAP-23 is a negative regulator of SNARE complex assembly and exosome secretion. OGT knockdown/overexpression, O-GlcNAcylation site analysis, SNARE complex co-IP, exosome quantification, cisplatin efflux assay Cell death discovery Medium 34001861
2022 ICAM-1 clustering in endothelial cells recruits SNAP23 (along with syntaxin-3 and -4) to mediate local release of endothelial chemokines (CXCL1/8/10); SNAP23 silencing shifts CTL transmigration from transcellular to paracellular route. siRNA silencing of SNAP23, immunofluorescence of ICAM-1/SNAP23 colocalization, T lymphocyte transmigration assay, chemokine release assay Cell reports Medium 35045291

Source papers

Stage 0 corpus · 100 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
1996 Identification of a novel syntaxin- and synaptobrevin/VAMP-binding protein, SNAP-23, expressed in non-neuronal tissues. The Journal of biological chemistry 294 8663154
2019 Long non-coding RNA HOTAIR promotes exosome secretion by regulating RAB35 and SNAP23 in hepatocellular carcinoma. Molecular cancer 230 30943982
1998 Relocation of the t-SNARE SNAP-23 from lamellipodia-like cell surface projections regulates compound exocytosis in mast cells. Cell 220 9727496
2000 Molecular mechanisms of platelet exocytosis: role of SNAP-23 and syntaxin 2 in dense core granule release. Blood 132 10648404
2008 Phosphorylation of SNAP-23 by IkappaB kinase 2 regulates mast cell degranulation. Cell 120 18692471
2000 Involvement of SNAP-23 and syntaxin 6 in human neutrophil exocytosis. Blood 115 11001914
1999 Proteins of the exocytotic core complex mediate platelet alpha-granule secretion. Roles of vesicle-associated membrane protein, SNAP-23, and syntaxin 4. The Journal of biological chemistry 112 9891020
2010 A neuronal role for SNAP-23 in postsynaptic glutamate receptor trafficking. Nature neuroscience 110 20118925
1998 Binary interactions of the SNARE proteins syntaxin-4, SNAP23, and VAMP-2 and their regulation by phosphorylation. Biochemistry 109 9693005
2009 Lipid droplets interact with mitochondria using SNAP23. Cell biology international 104 19524684
2000 Role of SNAP23 in insulin-induced translocation of GLUT4 in 3T3-L1 adipocytes. Mediation of complex formation between syntaxin4 and VAMP2. The Journal of biological chemistry 100 10713150
2004 Phosphorylation of SNAP-23 regulates exocytosis from mast cells. The Journal of biological chemistry 97 15611044
1999 Protein kinase B stimulates the translocation of GLUT4 but not GLUT1 or transferrin receptors in 3T3-L1 adipocytes by a pathway involving SNAP-23, synaptobrevin-2, and/or cellubrevin. The Journal of biological chemistry 97 10497159
2009 VAMP3, syntaxin-13 and SNAP23 are involved in secretion of matrix metalloproteinases, degradation of the extracellular matrix and cell invasion. Journal of cell science 96 19910495
2000 Molecular mechanisms of platelet exocytosis: role of SNAP-23 and syntaxin 2 and 4 in lysosome release. Blood 95 10961877
1999 Cultured glial cells express the SNAP-25 analogue SNAP-23. Glia 94 10417817
1997 SNAP-23 is not cleaved by botulinum neurotoxin E and can replace SNAP-25 in the process of insulin secretion. The Journal of biological chemistry 92 9407084
1999 Syntaxin 11 is associated with SNAP-23 on late endosomes and the trans-Golgi network. Journal of cell science 91 10036234
2004 The SNARE proteins SNAP-25 and SNAP-23 display different affinities for lipid rafts in PC12 cells. Regulation by distinct cysteine-rich domains. The Journal of biological chemistry 86 15542596
1999 SNAP-23 and SNAP-25 are palmitoylated in vivo. Biochemical and biophysical research communications 84 10329400
2013 IκB kinase phosphorylation of SNAP-23 controls platelet secretion. Blood 83 23613522
1997 Inhibition of the binding of SNAP-23 to syntaxin 4 by Munc18c. Biochemical and biophysical research communications 83 9168999
2005 Cholesterol-dependent syntaxin-4 and SNAP-23 clustering regulates caveolar fusion with the endothelial plasma membrane. The Journal of biological chemistry 74 16118213
2003 Tomosyn interacts with the t-SNAREs syntaxin4 and SNAP23 and plays a role in insulin-stimulated GLUT4 translocation. The Journal of biological chemistry 73 12832401
2003 Phosphorylation of SNAP-23 in activated human platelets. The Journal of biological chemistry 72 12930825
2003 Identification and characterization of Snapin as a ubiquitously expressed SNARE-binding protein that interacts with SNAP23 in non-neuronal cells. The Biochemical journal 71 12877659
2014 SNAP23, Syntaxin4, and vesicle-associated membrane protein 7 (VAMP7) mediate trafficking of membrane type 1-matrix metalloproteinase (MT1-MMP) during invadopodium formation and tumor cell invasion. Molecular biology of the cell 67 24807903
2000 Vimentin filaments in fibroblasts are a reservoir for SNAP23, a component of the membrane fusion machinery. Molecular biology of the cell 63 11029050
1998 SNAP-23 in rat kidney: colocalization with aquaporin-2 in collecting duct vesicles. The American journal of physiology 61 9815132
1997 Tissue distribution of SNAP-23 and its subcellular localization in 3T3-L1 cells. Biochemical and biophysical research communications 61 9020061
2002 Subcellular distribution of 3 functional platelet SNARE proteins: human cellubrevin, SNAP-23, and syntaxin 2. Blood 60 12010801
1998 SNAP-23 requirement for transferrin recycling in Streptolysin-O-permeabilized Madin-Darby canine kidney cells. The Journal of biological chemistry 57 9651373
2013 Morphine prevents lipopolysaccharide-induced TNF secretion in mast cells blocking IκB kinase activation and SNAP-23 phosphorylation: correlation with the formation of a β-arrestin/TRAF6 complex. Journal of immunology (Baltimore, Md. : 1950) 52 23960234
2014 Synaptotagmin-7 is an asynchronous calcium sensor for synaptic transmission in neurons expressing SNAP-23. PloS one 51 25422940
1999 SNAP23 promotes insulin-dependent glucose uptake in 3T3-L1 adipocytes: possible interaction with cytoskeleton. The American journal of physiology 50 10329959
2011 SNAP-23 and syntaxin-3 are required for chemokine release by mature human mast cells. Molecular immunology 48 21981832
1997 Botulinum neurotoxin B inhibits insulin-stimulated glucose uptake into 3T3-L1 adipocytes and cleaves cellubrevin unlike type A toxin which failed to proteolyze the SNAP-23 present. Biochemistry 47 9153412
2001 The last exon of SNAP-23 regulates granule exocytosis from mast cells. The Journal of biological chemistry 46 11350976
1999 Differential expression of SNAP-25 isoforms and SNAP-23 in the adrenal gland. Journal of neurochemistry 46 9886089
1997 Identification of two isoforms of the vesicle-membrane fusion protein SNAP-23 in human neutrophils and HL-60 cells. Biochemical and biophysical research communications 46 9070898
2004 SNAP-23 functions in docking/fusion of granules at low Ca2+. Molecular biology of the cell 45 14742706
2002 The heavy chain of conventional kinesin interacts with the SNARE proteins SNAP25 and SNAP23. Biochemistry 45 12475239
2017 VAMP3 and SNAP23 mediate the disturbed flow-induced endothelial microRNA secretion and smooth muscle hyperplasia. Proceedings of the National Academy of Sciences of the United States of America 44 28716920
2015 SNAP23 Regulates Endothelial Exocytosis of von Willebrand Factor. PloS one 44 26266817
1999 Organization of the secretory machinery in the rodent brain: distribution of the t-SNAREs, SNAP-25 and SNAP-23. Brain research 44 10411979
2021 Reduced O-GlcNAcylation of SNAP-23 promotes cisplatin resistance by inducing exosome secretion in ovarian cancer. Cell death discovery 39 34001861
2006 Potential involvement of galectin-3 and SNAP23 in Aeromonas hydrophila cytotoxic enterotoxin-induced host cell apoptosis. Microbial pathogenesis 38 16426811
2020 Small extracellular vesicles containing miR-30a-3p attenuate the migration and invasion of hepatocellular carcinoma by targeting SNAP23 gene. Oncogene 37 33110233
2007 Physical and functional interactions of SNAP-23 with annexin A2. American journal of respiratory cell and molecular biology 37 17575076
2004 Syntaxin 2 and SNAP-23 are required for regulated surfactant secretion. Biochemistry 37 15035620
2002 SNAP-23 is a target for calpain cleavage in activated platelets. The Journal of biological chemistry 37 12121992
2012 SNAP-23 regulates phagosome formation and maturation in macrophages. Molecular biology of the cell 36 23087210
2001 Cholecystokinin-regulated exocytosis in rat pancreatic acinar cells is inhibited by a C-terminus truncated mutant of SNAP-23. Pancreas 36 11484914
1999 SNAP-23 participates in SNARE complex assembly in rat adipose cells. The Biochemical journal 36 10051443
2010 The SNARE protein SNAP23 and the SNARE-interacting protein Munc18c in human skeletal muscle are implicated in insulin resistance/type 2 diabetes. Diabetes 35 20460426
2009 The t-SNAREs syntaxin4 and SNAP23 but not v-SNARE VAMP2 are indispensable to tether GLUT4 vesicles at the plasma membrane in adipocyte. Biochemical and biophysical research communications 35 20006577
2001 Role of SNAP-23 in trafficking of H+-ATPase in cultured inner medullary collecting duct cells. American journal of physiology. Cell physiology 35 11245593
2009 Single molecule measurements of mechanical interactions within ternary SNARE complexes and dynamics of their disassembly: SNAP25 vs. SNAP23. The Journal of physiology 34 19273577
2020 Botulinum toxin type A alleviates neuropathic pain and suppresses inflammatory cytokines release from microglia by targeting TLR2/MyD88 and SNAP23. Cell & bioscience 33 33298171
2011 Deletion of SNAP-23 results in pre-implantation embryonic lethality in mice. PloS one 33 21479242
2002 Function of the t-SNARE SNAP-23 and secretory carrier membrane proteins (SCAMPs) in exocytosis in mast cells. Molecular immunology 33 12217404
2021 Let-7a regulates EV secretion and mitochondrial oxidative phosphorylation by targeting SNAP23 in colorectal cancer. Journal of experimental & clinical cancer research : CR 32 33446221
2011 VAMP3 regulates podosome organisation in macrophages and together with Stx4/SNAP23 mediates adhesion, cell spreading and persistent migration. Experimental cell research 32 21586284
2005 Identification of soluble N-ethylmaleimide-sensitive factor attachment protein receptor exocytotic machinery in human plasma cells: SNAP-23 is essential for antibody secretion. Journal of immunology (Baltimore, Md. : 1950) 31 16272324
2016 Septin 7 reduces nonmuscle myosin IIA activity in the SNAP23 complex and hinders GLUT4 storage vesicle docking and fusion. Experimental cell research 30 28011197
2013 Plin2 inhibits cellular glucose uptake through interactions with SNAP23, a SNARE complex protein. PloS one 30 24040030
2005 Direct interaction between SNAP-23 and L-type Ca2+ channel. Journal of cellular and molecular medicine 28 15963257
2020 Pancreas-specific SNAP23 depletion prevents pancreatitis by attenuating pathological basolateral exocytosis and formation of trypsin-activating autolysosomes. Autophagy 26 33213278
2006 A dominant-negative variant of SNAP-23 decreases the cell surface expression of the neuronal glutamate transporter EAAC1 by slowing constitutive delivery. Neurochemistry international 26 16516346
2019 SNAP23 is required for constitutive and regulated exocytosis in mouse oocytes†. Biology of reproduction 25 31201423
2016 Opposing roles for SNAP23 in secretion in exocrine and endocrine pancreatic cells. The Journal of cell biology 25 27697926
2013 SNAP-23 and VAMP-3 contribute to the release of IL-6 and TNFα from a human synovial sarcoma cell line. The FEBS journal 25 24373201
1999 Phosphorylation of SNAP-23 by the novel kinase SNAK regulates t-SNARE complex assembly. Molecular biology of the cell 24 10588641
2015 Identification and Characterization of Botulinum Neurotoxin A Substrate Binding Pockets and Their Re-Engineering for Human SNAP-23. Journal of molecular biology 23 26523682
2018 SNAP23 regulates BAX-dependent adipocyte programmed cell death independently of canonical macroautophagy. The Journal of clinical investigation 22 30102258
2017 Electrostatic anchoring precedes stable membrane attachment of SNAP25/SNAP23 to the plasma membrane. eLife 22 28240595
2017 Rab5 is critical for SNAP23 regulated granule-granule fusion during compound exocytosis. Scientific reports 22 29127297
2015 SNAP23 is selectively expressed in airway secretory cells and mediates baseline and stimulated mucin secretion. Bioscience reports 22 26182382
2012 Annexin A7 and SNAP23 interactions in alveolar type II cells and in vitro: a role for Ca(2+) and PKC. Biochimica et biophysica acta 22 22713544
2009 Distribution of the SNAP25 and SNAP23 synaptosomal-associated protein isoforms in rat cerebellar cortex. Neuroscience 22 19735702
1999 Intracellular localization of SNAP-23 to endosomal compartments. Biochemical and biophysical research communications 22 10049710
2022 The endothelial diapedesis synapse regulates transcellular migration of human T lymphocytes in a CX3CL1- and SNAP23-dependent manner. Cell reports 21 35045291
2019 Syntaxin 11 regulates the stimulus-dependent transport of Toll-like receptor 4 to the plasma membrane by cooperating with SNAP-23 in macrophages. Molecular biology of the cell 21 30811271
2003 SNARE protein degradation upon platelet activation: calpain cleaves SNAP-23. Journal of cellular physiology 21 12494459
1997 Botulinum E toxin light chain does not cleave SNAP-23 and only partially impairs insulin stimulation of GLUT4 translocation in 3T3-L1 cells. Biochemical and biophysical research communications 21 9268721
2015 SNAP23/25 and VAMP2 mediate exocytic event of transferrin receptor-containing recycling vesicles. Biology open 20 26092867
2016 Immunofluorescence microscopy of SNAP23 in human skeletal muscle reveals colocalization with plasma membrane, lipid droplets, and mitochondria. Physiological reports 19 26733245
2016 SNAP23 promotes the malignant process of ovarian cancer. Journal of ovarian research 18 27855700
2020 Regulatory Mechanism of SNAP23 in Phagosome Formation and Maturation. Yonago acta medica 17 32884432
2017 Phosphorylation of SNAP-23 regulates its dynamic membrane association during mast cell exocytosis. Biology open 17 28784843
2020 SNAP23 depletion enables more SNAP25/calcium channel excitosome formation to increase insulin exocytosis in type 2 diabetes. JCI insight 16 32051343
2018 Platelet-specific deletion of SNAP23 ablates granule secretion, substantially inhibiting arterial and venous thrombosis in mice. Blood advances 16 30573565
2016 Expression of genes involved in lipid droplet formation (BSCL2, SNAP23 and COPA) during porcine in vitro adipogenesis. Journal of applied genetics 16 27108337
2015 Expression of the SNARE protein SNAP-23 is essential for cell survival. PloS one 16 25706117
2007 SNAP-23 and syntaxin-2 localize to the extracellular surface of the platelet plasma membrane. Blood 16 17485553
2005 Mutations of syntaxin 11 and SNAP23 genes as causes of familial hemophagocytic lymphohistiocytosis were not found in Japanese people. Journal of human genetics 16 16180048
2007 SNAP-23 is not essential for constitutive exocytosis in HeLa cells. FEBS letters 15 17825825
2018 SNAP23-Kif5 complex controls mGlu1 receptor trafficking. Journal of molecular cell biology 14 29762713
2012 Role of the SNARE protein SNAP23 on cAMP-stimulated renin release in mouse juxtaglomerular cells. American journal of physiology. Renal physiology 14 23269646
2008 Heterogeneous expression of SNARE proteins SNAP-23, SNAP-25, Syntaxin1 and VAMP in human parathyroid tissue. Molecular and cellular endocrinology 14 18457912

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