Affinage

MIB2

E3 ubiquitin-protein ligase MIB2 · UniProt Q96AX9

Audit flag: model fault
Length
955 aa
Mass
103.7 kDa
Annotated
2026-06-13
26 papers in source corpus 21 papers cited in narrative 21 extracted findings

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

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Mechanism profile

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Evidence

Reading pass · 21 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2003 MIB2 (skeletrophin) binds actin monomer, as demonstrated by yeast two-hybrid screening and co-immunoprecipitation experiments. Yeast two-hybrid screen; co-immunoprecipitation The American journal of pathology Medium 14507647
2005 MIB2 (skeletrophin) is a RING finger-dependent E3 ubiquitin ligase that binds the intracellular region of Notch ligand Jagged-2 (but not Delta-1, -3, -4, or Jagged-1) and ubiquitinates it; RING-mutated skeletrophin lacks this activity. Exogenous skeletrophin (but not RING mutant) in Jagged-2-positive myeloma cells induced Hes-1 expression in stromal cells via Notch signaling. In vitro autoubiquitination assay with recombinant proteins; RING domain mutagenesis; cell-based functional assay The American journal of pathology High 15920166
2006 MIB2 (skeletrophin) reduces melanoma cell invasion in vitro and in vivo and suppresses colony formation in soft agar in a RING motif-dependent manner; it also downregulates Met oncogene transcription and increases Hes1 transcription. The RING mutant lacks these activities. Exogenous expression; RING domain mutagenesis; invasion assays; soft agar colony formation; in vivo xenograft Oncogene Medium 16715130
2006 Zebrafish Mib2 has a C-terminal-most RING finger-dependent E3 ubiquitin ligase activity; Mib and Mib2 are reciprocal E3 ubiquitin ligases and substrates of each other; they share DeltaC as a common substrate but differ in DeltaD internalization; Mib2 binds differently from Mib to extracellular and intracellular parts of DeltaA and DeltaC. In vitro ubiquitination assay; RING domain mutagenesis; Delta internalization assay; binding assays Journal of molecular biology High 17196985
2007 Zebrafish Mib2 co-localizes with Mib in transfected cells and functions redundantly with Mib in regulating Notch signaling in embryos; dominant-negative Mib alleles compete with Mib2 in DeltaC ubiquitylation and internalization. Co-localization in transfected cells; genetic epistasis in zebrafish embryos; ubiquitylation/internalization assay Developmental biology Medium 17331493
2007 Targeted disruption of Mib2 in mice causes failure to close the cranial neural tube (exencephaly) with variable penetrance depending on genetic background, indicating a role in neural tube closure but not complete developmental essentiality. Targeted gene knockout in mice; developmental phenotype analysis Genesis (New York, N.Y. : 2000) Medium 17987667
2011 MIB2 is a component of the activated BCL10 signaling complex; it directly interacts with BCL10 (shown by in vitro translation and pulldown); MIB2 promotes autoubiquitination and ubiquitination of IKKγ/NEMO and recruits/activates TAK1, thereby controlling BCL10-dependent NF-κB activation. MIB2 knockdown inhibited BCL10-dependent NF-κB activation. Proteomics identification; in vitro pulldown; overexpression; RNAi knockdown; NF-κB reporter assay The Journal of biological chemistry Medium 21896478
2014 CYLD (ubiquitin hydrolase) interacts with MIB2; co-expression of CYLD and MIB2 stabilizes MIB2 protein levels and is associated with reduced levels of JAG2; silencing CYLD increases JAG2 expression and upregulates Notch signaling. Proteomics/co-IP interaction screen; co-expression experiments; siRNA knockdown; Notch target gene expression Oncotarget Medium 25565632
2017 A missense variant MIB2 p.V742G shows reduced ubiquitination activity in vitro; expression of MIB2 variants (p.V742G and p.V984L) affects NOTCH signaling, proliferation, and apoptosis in primary rat cardiomyocytes. In vitro ubiquitination assay; functional cell-based assay in primary cardiomyocytes; whole exome sequencing with family segregation Human molecular genetics Medium 28013292
2017 In Drosophila, the ankyrin repeat domains (in entirety) and Mib-specific (MIB) domains of Mib2 are required for its major function in maintaining larval skeletal and visceral muscle integrity; the RING finger domain is required for adult flight muscle development. Missense mutations in the MIB domain and RING finger produce viable but flightless hypomorphic phenotypes. Domain deletion analysis; forward genetic screen; in vivo imaging; functional dissection with novel alleles PloS one Medium 28282454
2018 MIB2 catalyzes Lys-63-linked ubiquitination of GABAB1 at multiple sites, sorting GABAB receptors to lysosomes for degradation. CaMKIIβ (but not CaMKIIα) promotes this MIB2-mediated Lys-63-linked ubiquitination; phosphorylation of GABAB1 at Ser-867 by CaMKII is required, as the phosphomimetic S867D mutant shows strongly increased Lys-63-linked ubiquitination and reduced surface expression, while inactivation of Ser-867 reduces ubiquitination. Site-directed mutagenesis; overexpression/knockdown; ubiquitin linkage-specific assays; surface receptor quantification in cortical neurons Molecular neurobiology High 29881949
2019 MIB2 promotes proteasomal degradation of CYLD via Lys-48-linked polyubiquitination at Lys-338 and Lys-530 of CYLD; the ankyrin repeat in MIB2 interacts with the third CAP domain of CYLD. MIB2-dependent CYLD degradation activates NF-κB signaling via TNFα stimulation and LUBAC. Mib2-knockout mice show reduced IL-6, reduced arthritic inflammation, and suppressed NF-κB signaling. AlphaScreen; pulldown assay; immunofluorescence; Mib2 KO mice; ubiquitination mapping (K48-linked, site-specific) The Journal of biological chemistry High 31366726
2021 Gm364 (a transmembrane protein) directly binds and anchors MIB2 on the membrane; membrane-localized MIB2 ubiquitinates and activates DLL3, which binds and activates Notch2; the resulting NICD2 directly activates AKT in the cytoplasm to regulate oocyte meiosis and quality. Genetic knockout; co-IP/binding assays; ubiquitination assay; Notch pathway activation assay; oocyte developmental phenotype Cell death and differentiation Medium 34635817
2023 MIB2 is required for PD-L1 transport from the trans-Golgi network to the plasma membrane; mechanistically, MIB2 catalyzes nonproteolytic K63-linked ubiquitination of PD-L1, facilitating RAB8-mediated exocytosis to the plasma membrane. MIB2 deficiency reduces surface PD-L1 and promotes antitumor T cell immunity in mice. MIB2 knockdown/knockout; ubiquitination assay (K63-linkage specific); RAB8 interaction/trafficking assay; mouse tumor models; surface PD-L1 quantification The Journal of clinical investigation High 36719382
2023 FAT1 (protocadherin) interacts with MIB2 E3 ubiquitin ligase; MIB2 mediates FAT1-induced ubiquitination and degradation of YAP/TAZ in endothelial cells. Loss of MIB2 reduces YAP/TAZ degradation, increases YAP/TAZ protein levels and target gene expression, and increases endothelial cell proliferation and angiogenesis in vivo. Co-IP/interactome analysis; MIB2 knockdown in vitro and in vivo; YAP/TAZ ubiquitination assay; angiogenesis models Nature communications High 37031213
2023 MIB2 interacts with CARD6 and promotes K48-linked CARD6 polyubiquitination and proteasomal degradation; high fructose upregulates MIB2 expression, leading to CARD6 degradation and consequent ASK1/JNK1/2 activation and lipid accumulation in hepatocytes. MIB2 siRNA reverses these effects. Immunoprecipitation; immunofluorescence; siRNA knockdown; immunoblotting Food & function Medium 37186242
2024 MIB2 expression is increased in SN (surrounded nucleolus) oocytes; specific depletion of MIB2 in SN oocytes disrupts meiotic apparatus and causes aneuploidy, impaired meiotic maturation, and defective early embryo development. Overexpression of MIB2 in NSN oocytes facilitates chromatin configuration transition from NSN to SN and reduces spindle/chromosome disorganization. Quantitative proteomics; MIB2-specific depletion; overexpression; meiotic phenotype analysis; aneuploidy quantification Molecular & cellular proteomics : MCP Medium 39019259
2025 Ebola virus VP35 contains an NNLNS motif (residues 201–205) that serves as a direct binding site for MIB2; this motif is critical for VP35-dependent inhibition of MIB2-mediated interferon induction. MIB2 binding to VP35 also inhibits EBOV minigenome (RNA synthesis) activity. Mutational analysis of NNLNS motif; protein-protein interaction assays; IFN induction assay; EBOV minigenome assay Proceedings of the National Academy of Sciences of the United States of America Medium 40982696
2025 MIB2 directly interacts with Runx2 transcription factor, ubiquitinating and degrading Runx2, which inhibits Hmgcs2 transcription and impedes fatty acid metabolic processes in cardiomyocytes. Cardiac-specific MIB2 overexpression in HFpEF mice worsens cardiac dysfunction and lipid accumulation. Immunoprecipitation; dual luciferase reporter assay; proteomic analysis; AAV9-mediated cardiac overexpression in mice Journal of cellular and molecular medicine Medium 40159625
2025 MIB2 physically interacts with SUZ12 (PRC2 component) via its MIB/HERC and ZZ-type domains; MIB2 mediates ubiquitination of SUZ12, promoting its degradation. MIB2 knockdown reduces SUZ12 protein levels (despite increased ubiquitination), decreases H3K27me3, upregulates PRC2 target genes, and reduces proliferation/increases apoptosis in PNH clone cells. Immunoprecipitation and mass spectrometry; domain mapping; RNA-seq; western blotting; flow cytometry; colony formation assay Journal of cellular and molecular medicine Medium 40478202
2025 Loss of FAT1 in tumor cells (including HNSCC) leads to decreased YAP/TAZ ubiquitination and degradation; MIB2 is identified as an interaction partner of the intracellular part of FAT1 by interactome analysis; MIB2 suppression recapitulates FAT1 loss (increased YAP/TAZ levels, target gene expression, proliferation in vitro, and tumor growth in vivo). Interactome analysis of FAT1 cytoplasmic domain; siRNA knockdown; ubiquitination assay; in vivo xenograft PloS one Medium 40478800

Source papers

Stage 0 corpus · 26 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2022 MIB2: metal ion-binding site prediction and modeling server. Bioinformatics (Oxford, England) 94 35904542
2023 PD-L1 translocation to the plasma membrane enables tumor immune evasion through MIB2 ubiquitination. The Journal of clinical investigation 52 36719382
2005 Skeletrophin, a novel ubiquitin ligase to the intracellular region of Jagged-2, is aberrantly expressed in multiple myeloma. The American journal of pathology 49 15920166
2007 The characterization of zebrafish antimorphic mib alleles reveals that Mib and Mind bomb-2 (Mib2) function redundantly. Developmental biology 45 17331493
2011 The E3 ubiquitin ligase mind bomb-2 (MIB2) protein controls B-cell CLL/lymphoma 10 (BCL10)-dependent NF-κB activation. The Journal of biological chemistry 41 21896478
2023 Endothelial FAT1 inhibits angiogenesis by controlling YAP/TAZ protein degradation via E3 ligase MIB2. Nature communications 35 37031213
2006 A ubiquitin ligase, skeletrophin, is a negative regulator of melanoma invasion. Oncogene 34 16715130
2006 Zebrafish Mib and Mib2 are mutual E3 ubiquitin ligases with common and specific delta substrates. Journal of molecular biology 34 17196985
2019 The E3 ubiquitin ligase MIB2 enhances inflammation by degrading the deubiquitinating enzyme CYLD. The Journal of biological chemistry 29 31366726
2014 The cylindromatosis gene product, CYLD, interacts with MIB2 to regulate notch signalling. Oncotarget 26 25565632
2018 Ca2+/Calmodulin-Dependent Protein Kinase II (CaMKII) β-Dependent Phosphorylation of GABAB1 Triggers Lysosomal Degradation of GABAB Receptors via Mind Bomb-2 (MIB2)-Mediated Lys-63-Linked Ubiquitination. Molecular neurobiology 24 29881949
2003 Down-regulation of a novel actin-binding molecule, skeletrophin, in malignant melanoma. The American journal of pathology 24 14507647
2021 Gm364 coordinates MIB2/DLL3/Notch2 to regulate female fertility through AKT activation. Cell death and differentiation 14 34635817
2007 Targeted disruption of Mib2 causes exencephaly with a variable penetrance. Genesis (New York, N.Y. : 2000) 13 17987667
2023 Nuciferine blocks MIB2-mediated CARD6 polyubiquitination and degradation in the amelioration of high fructose-induced liver lipid accumulation. Food & function 8 37186242
2017 MIB2 variants altering NOTCH signalling result in left ventricle hypertrabeculation/non-compaction and are associated with Ménétrier-like gastropathy. Human molecular genetics 8 28013292
2017 Identification of the essential protein domains for Mib2 function during the development of the Drosophila larval musculature and adult flight muscles. PloS one 7 28282454
2005 Skeletrophin, a novel RING molecule controlled by the chromatin remodeling complex, is downregulated in malignant melanoma. DNA and cell biology 7 15869411
2025 Taraxerol induces ferroptosis in breast cancer by targeting Nrf2 transcriptional activity to promote MIB2-mediated GPX4 ubiquitination. Phytomedicine : international journal of phytotherapy and phytopharmacology 6 40592077
2025 Mib2 Regulates Lipid Metabolism in Heart Failure With Preserved Ejection Fraction via the Runx2-Hmgcs2 Axis. Journal of cellular and molecular medicine 3 40159625
2025 The tumor suppressor FAT1 controls YAP/TAZ protein degradation and tumor cell proliferation through E3 ligase MIB2. PloS one 2 40478800
2025 Ebola virus VP35 NNLNS motif modulates viral RNA synthesis and MIB2-mediated signaling. Proceedings of the National Academy of Sciences of the United States of America 2 40982696
2025 Ebola virus VP35 NNLNS motif modulates viral RNA synthesis and MIB2-mediated signaling. bioRxiv : the preprint server for biology 1 40777377
2023 MIB2 promotes the progression of non-small cell lung cancer by regulating cell cycle control pathways. Genes & genomics 1 37436668
2025 MIB2-Mediated SUZ12 Ubiquitination Regulates Clonal Proliferation in Patients With Paroxysmal Nocturnal Haemoglobinuria. Journal of cellular and molecular medicine 0 40478202
2024 MIB2 Functions in Oocyte Meiosis by Modulating Chromatin Configuration. Molecular & cellular proteomics : MCP 0 39019259

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