Affinage

USP15

Ubiquitin carboxyl-terminal hydrolase 15 · UniProt Q9Y4E8

Length
981 aa
Mass
112.4 kDa
Annotated
2026-06-10
100 papers in source corpus 50 papers cited in narrative 50 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 8/8 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

USP15 is a cysteine deubiquitylase of the USP family that functions as a reversible-ubiquitylation switch across signaling, genome-maintenance, immune, and RNA-processing pathways by cleaving ubiquitin from a broad set of substrates (PMID:10444327, PMID:22344298). Its catalytic core adopts a canonical USP fold whose catalytic triad can toggle between aligned (active) and misaligned (inactive) configurations, and an N-terminal DUSP-UBL module forms an elongated tandem arrangement that mediates substrate-targeting interactions (PMID:21848306, PMID:30228188, PMID:35605756); a zinc-finger motif is required for poly-ubiquitin chain cleavage (PMID:16005295). Substrate selection and nuclear/cytoplasmic distribution are governed by adaptor binding and phosphorylation: the spliceosome recycling factor SART3 recruits USP15 in a bipartite mode to deliver it to nuclear substrates, while CDK-type phosphorylation at Thr149/Thr219 and isoform-specific events control SART3 association and nuclear translocation (PMID:27255711, PMID:31330151, PMID:29429988). In TGF-β/BMP signaling USP15 stabilizes the type I receptors TβR-I and ALK3/BMPR1A and acts on SMURF2 and receptor-activated SMADs to potentiate pathway output (PMID:21947082, PMID:22344298, PMID:24850914, PMID:26435193). It controls innate immunity bidirectionally, stabilizing TRIM25 to enhance RIG-I-dependent type I IFN yet removing activating ubiquitin from TBK1, cGAS, and CARD9, and deubiquitylating IκBα and TAB2/TAB3 to tune NF-κB (PMID:24399297, PMID:27721430, PMID:32814047, PMID:33093067, PMID:36243958, PMID:31477895, PMID:31903660). At DNA double-strand breaks phospho-Ser678 USP15 is recruited by MDC1 to deubiquitylate BARD1 and promote homologous recombination, and it stabilizes TOP2A and PARP1 to safeguard genomic stability, with USP15-null mice showing genomic instability (PMID:30874560, PMID:29429988, PMID:37012401). USP15 additionally regulates oxidative-stress (Keap1/Nrf2) and p53 axes via MDM2, opposes Parkin-driven mitophagy, and participates in spliceosomal deubiquitylation of PRP31/PRP3 together with SART3 and USP15's paralog USP4 (PMID:23727018, PMID:24777531, PMID:24852371, PMID:28088760). Mouse viability requires a functional copy of either USP15 or its paralog USP4, indicating genetic redundancy (PMID:26503449).

Mechanistic history

Synthesis pass · year-by-year structured walk · 16 steps
  1. 1999 High

    Established that USP15 is a bona fide ubiquitin-specific protease, defining its core enzymatic identity.

    Evidence in vitro deubiquitinase assay with recombinant protein against linear ubiquitin fusion substrates

    PMID:10444327

    Open questions at the time
    • No physiological substrate identified
    • Chain-linkage preference not addressed
    • No structural basis for catalysis
  2. 2005 High

    Linked USP15 to the COP9 signalosome and showed a zinc-finger motif is required for poly-ubiquitin chain cleavage and for stabilizing an E3 ligase, moving beyond minimal substrates to E3-protective roles.

    Evidence co-purification, tetra-Ub pulldown, Zn-finger mutagenesis, cotransfection ubiquitination assay (Rbx1)

    PMID:16005295

    Open questions at the time
    • Endogenous CSN-dependent substrate repertoire unmapped
    • Stoichiometry within CSN unknown
  3. 2011 High

    Defined USP15 as a positive regulator of TGF-β/BMP signaling by reversing R-SMAD monoubiquitylation required for promoter occupancy, and resolved the elongated DUSP-UBL architecture.

    Evidence in vitro deubiquitylation, ChIP, knockdown in cells and Xenopus; X-ray/SAXS of DUSP-UBL

    PMID:21848306 PMID:21947082

    Open questions at the time
    • How DUSP-UBL selects SMAD substrates unresolved
    • Catalytic-domain structure not yet solved
  4. 2012 High

    Showed USP15 acts at the receptor level of TGF-β signaling by deubiquitylating TβR-I via the SMAD7-SMURF2 complex, with in vivo relevance to glioblastoma.

    Evidence RNAi screen, Co-IP, in vitro deubiquitylation, patient-derived orthotopic mouse model; plus BRAP/MAPK study

    PMID:22344298 PMID:23105109

    Open questions at the time
    • Direct vs SMURF2-mediated effect on receptor not fully separated
    • Selectivity among receptor-complex components unclear
  5. 2013 High

    Expanded USP15 substrates into oxidative-stress and transcription control, deubiquitylating Keap1 to suppress Nrf2 and stabilizing newly synthesized REST.

    Evidence in vitro deubiquitylation, Co-IP, reporter assays, polysome fractionation, cycloheximide chase

    PMID:23708518 PMID:23727018

    Open questions at the time
    • Mechanism coupling USP15 to translation/polysomes unknown
    • Cell-type specificity of Keap1 effect not defined
  6. 2014 High

    Demonstrated broad regulatory reach — stabilizing MDM2 to restrain T-cell/p53 responses, opposing Parkin mitophagy, stabilizing TRIM25 for IFN, deubiquitylating histone H2B via SART3, and targeting ALK3 in BMP signaling — establishing USP15 as a multi-pathway DUB.

    Evidence in vitro deubiquitylation, KO mice, mitophagy and patient-fibroblast assays, MS, SART3 in vitro assays, Xenopus

    PMID:24399297 PMID:24526689 PMID:24777531 PMID:24850914 PMID:24852371

    Open questions at the time
    • Determinants of substrate choice across pathways unknown
    • Whether one localization pool serves all substrates unclear
  7. 2015 Medium

    Refined the TGF-β mechanism by identifying SMURF2 Lys734 deubiquitylation, and established USP4/USP15 genetic redundancy required for mouse viability.

    Evidence proteomic MS, site-directed mutagenesis, in vitro deubiquitylation; Usp4/Usp15 mouse crosses

    PMID:26435193 PMID:26503449

    Open questions at the time
    • Degree of substrate overlap with USP4 not catalogued
    • Tissue-specific redundancy unresolved
  8. 2016 High

    Connected USP15 activity to physiology and disease in vivo, showing a hypomorphic mutation dampens type I IFN and protects against neuroinflammation, and that TGF-β drives USP15 translation to stabilize p53.

    Evidence ENU mouse model, RNA-seq, neuroinflammation models; Co-IP, pathway inhibitors

    PMID:27721430 PMID:27893708

    Open questions at the time
    • Direct substrate driving IFN phenotype in vivo not pinpointed
    • Generality of TGF-β→mTOR→USP15 axis untested
  9. 2017 High

    Established the structural and regulatory logic of SART3-mediated targeting and isoform/phospho control, and defined spliceosomal substrates PRP31/PRP3.

    Evidence crystal structures with ITC/mutagenesis; isoform-specific pulldowns; in vitro deubiquitylation of PRP31/PRP3 with USP4/SART3

    PMID:27255711 PMID:28088760 PMID:28276505

    Open questions at the time
    • Functional consequence of exon-7 isoform switch in vivo limited
    • How splicing-substrate deubiquitylation integrates with cell cycle unclear
  10. 2018 High

    Resolved that the catalytic core has an intrinsically misaligned (inactive) triad with low monoubiquitin affinity, and placed USP15 directly in DNA-damage repair via MDC1-recruited, phospho-S678-dependent BARD1 deubiquitylation and TOP2A stabilization.

    Evidence X-ray crystallography, ITC, inhibition assay; Co-IP, phospho-mutant analysis, HR assays, KO mice; imaging and ubiquitination assays

    PMID:29429988 PMID:29593334 PMID:30228188 PMID:30874560

    Open questions at the time
    • What triggers triad activation on substrate not defined
    • Kinase responsible for S678/S229 phosphorylation in repair not identified
  11. 2019 High

    Defined inhibitor-accessible conformational control via ubiquitin variants, mapped phospho-regulated nuclear targeting through SART3, and uncovered non-canonical Hrd1 ubiquitination that inactivates USP15 toward IκBα.

    Evidence UbV engineering with crystal structures and cell assays; fractionation/MS phospho-mutant analysis; E3 screen, KO mice, sepsis model

    PMID:29299163 PMID:30713027 PMID:31330151 PMID:31477895

    Open questions at the time
    • In vivo kinase(s) for Thr149/Thr219 not established
    • Physiological scope of Hrd1-mediated USP15 inactivation unclear
  12. 2020 Medium

    Broadened immune and stem-cell roles, showing USP15 deubiquitylates TET2, CARD9, TBK1, and stabilizes TAB2/TAB3 and FUS, with TIFAB acting as an activator of USP15 toward MDM2/KEAP1.

    Evidence in vitro deubiquitylation, KO models, linkage-specific ubiquitination assays, proteomics, leukemia/transplantation models

    PMID:31903660 PMID:32101751 PMID:32814047 PMID:32839293 PMID:32948596 PMID:33093067 PMID:33378683

    Open questions at the time
    • How a single DUB achieves opposing IFN outcomes mechanistically unresolved
    • Effector-driven (TIFAB) activation generality untested
  13. 2021 Medium

    Extended USP15 to additional disease-relevant substrates including CRL4CRBN neosubstrates, mutant p53-R175H, ERα, BECN1, and TUT1, linking it to drug resistance, autophagy, and RNA metabolism.

    Evidence ubiquitination/deubiquitination assays, knockdown/KO, lenalidomide/tamoxifen sensitivity, KO mice with U6 snRNA readout

    PMID:29593334 PMID:32839293 PMID:33771975 PMID:34583995 PMID:35422093

    Open questions at the time
    • Catalytic vs scaffold contributions not separated for several substrates
    • Direct vs indirect deubiquitylation not always established
  14. 2022 Medium

    Showed USP15 promotes cGAS activation not only by deubiquitylation but by driving cGAS liquid condensation through its intrinsic disordered region, adding a phase-separation dimension to its function.

    Evidence Co-IP, in vitro deubiquitylation, LLPS assays, IDR deletion mutants, cGAS activation assays

    PMID:36243958

    Open questions at the time
    • Single-lab finding awaiting independent confirmation
    • Quantitative role of IDR vs catalysis in vivo unclear
  15. 2023 High

    Established the active/inactive triad switch as ligand-independent and consolidated genome-stability and tumor-immune roles via PARP1 stabilization (regulated by ER/PR/HER2), SMYD3-driven chemokine induction, neurofilament regulation opposing CRL3gigaxonin, and YAP1/TAZ stabilization.

    Evidence multiple crystal structures; Co-IP, deubiquitination assays, ChIP, degron mapping, tumor and disease models

    PMID:35605756 PMID:36635430 PMID:37012401 PMID:37903270 PMID:40323348

    Open questions at the time
    • What endogenously switches the triad state unresolved
    • Substrate-specific conformational requirements unmapped
  16. 2024 Medium

    Continued substrate expansion into cancer metabolism and drug sensitivity, deubiquitylating HKDC1 and antagonizing TRIM21 on ACSL4.

    Evidence Co-IP, ubiquitination assays, metabolic/glycolysis readouts, xenografts, clinical correlation

    PMID:38182686 PMID:39164728

    Open questions at the time
    • Direct binding vs indirect stabilization not fully resolved
    • In vivo significance beyond tumor models unknown

Open questions

Synthesis pass · forward-looking unresolved questions
  • What governs USP15 substrate selectivity and the active/inactive catalytic-triad switch across its many pathways remains the central open question.
  • No unifying model linking phosphorylation, SART3/adaptor binding, and triad activation to substrate choice
  • Endogenous trigger of triad alignment unknown
  • Relative weight of catalytic vs scaffold/IDR functions per substrate undefined

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0140096 catalytic activity, acting on a protein 8 GO:0016787 hydrolase activity 4 GO:0098772 molecular function regulator activity 3
Localization
GO:0005634 nucleus 4 GO:0005829 cytosol 2 GO:0005730 nucleolus 1 GO:0005739 mitochondrion 1
Pathway
R-HSA-168256 Immune System 5 R-HSA-162582 Signal Transduction 4 R-HSA-392499 Metabolism of proteins 3 R-HSA-73894 DNA Repair 3 R-HSA-8953854 Metabolism of RNA 3
Complex memberships
COP9 signalosome (CSN)USP15-SART3-USP4 spliceosomal complex

Evidence

Reading pass · 50 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
1999 Recombinant USP15 demonstrated ubiquitin-specific protease activity against engineered linear fusions of ubiquitin to beta-galactosidase and glutathione S-transferase, and can cleave the ubiquitin-proline bond. USP15 protein consists of 952 amino acids and contains conserved Cys and His boxes present in all UBP family members. In vitro deubiquitinase activity assay with recombinant protein against linear ubiquitin fusion substrates Genomics High 10444327
2005 USP15 co-purifies with the human COP9 signalosome (CSN) complex. A novel zinc finger in USP15 is essential for cleavage of poly-ubiquitin chains; mutation of a single conserved cysteine in the Zn-binding motif abolishes poly-Ub substrate degradation and reduces tetra-Ub binding. Wild-type but not Zn-finger-mutant USP15 stabilizes the E3 ligase Rbx1 by reversing its poly/autoubiquitination. Co-purification, pulldown with tetra-Ub, site-directed mutagenesis, cotransfection ubiquitination assay Current biology : CB High 16005295
2009 USP15 stabilizes APC (adenomatous polyposis coli) protein in the context of the COP9 signalosome-associated complex. Knockdown of USP15 or overexpression of a catalytically inactive USP15 mutant accelerates APC proteolysis, demonstrating that CSN-associated USP15 protects APC from ubiquitin-mediated degradation. siRNA knockdown, overexpression of catalytically inactive mutant, Western blot for APC stability Journal of molecular biology Medium 19576224
2009 USP15 interacts with HPV16 E6 protein (identified by tandem affinity purification). Overexpression of USP15 increases E6 protein levels and siRNA-mediated knockdown of USP15 decreases E6 protein levels, implicating USP15 in stabilization of E6 through deubiquitylation. Tandem affinity purification, siRNA knockdown, overexpression, Western blot Journal of virology Medium 19553310
2011 USP15 is a deubiquitylating enzyme for receptor-activated SMADs (R-SMADs: SMAD1, 2, 3). USP15 primarily opposes R-SMAD monoubiquitylation at DNA-binding domains, which normally prevents promoter recognition. USP15 is required for TGF-β and BMP responses in mammalian cells and Xenopus embryos and is necessary for SMAD complex occupancy of endogenous target promoters. siRNA knockdown, overexpression, ChIP, in vitro deubiquitylation assays, Xenopus embryo knockdown Nature cell biology High 21947082
2011 Crystal structure of the USP15 N-terminal DUSP and UBL domains at 1.5 Å resolution reveals an 80 Å elongated arrangement with the DU domains aligned in tandem, connected through a β-hairpin (DU finger) that forms an intricate hydrogen-bonding network. The UBL domain is monomeric in solution and unlikely to act as a ubiquitin mimic. X-ray crystallography, analytical ultracentrifugation, SAXS, gel filtration Biochemistry High 21848306
2012 USP15 binds to the SMAD7-SMURF2 complex and deubiquitinates and stabilizes type I TGF-β receptor (TβR-I), leading to enhanced TGF-β signaling. Identified via functional RNAi screen; downregulation or inhibition of USP15 decreases TGF-β activity in patient-derived orthotopic mouse glioblastoma models. Functional RNAi screen, Co-IP, in vitro deubiquitylation, orthotopic mouse model with patient-derived cells Nature medicine High 22344298
2012 USP15 interacts with the E3 ligase BRAP/IMP through the N-terminal DUSP-UBL domain of USP15 and the coiled-coil region of BRAP. USP15 opposes BRAP autoubiquitylation and stabilizes BRAP protein through its catalytic activity. USP15 depletion destabilizes BRAP via proteasomal degradation, reduces CRAF levels, and decreases MAPK signaling amplitude in response to EGF and PDGF. Co-IP, domain mapping, catalytically inactive mutant rescue, siRNA knockdown, Western blot, EGF/PDGF stimulation assays The Journal of biological chemistry Medium 23105109
2013 USP15 stabilizes newly synthesized REST (RE1-silencing transcription factor) by antagonizing its polyubiquitylation in a catalytic-activity-dependent manner. USP15 specifically promotes de novo REST synthesis (associates with polysomes) rather than protecting pre-existing REST, and is required for rapid accumulation of newly synthesized REST at mitotic exit. siRNA screening, polysome fractionation, cycloheximide chase, ubiquitination assays, cell cycle analysis Cell cycle (Georgetown, Tex.) Medium 23708518
2013 USP15 specifically deubiquitinates Keap1 (not Nrf2 directly), which promotes incorporation of deubiquitinated Keap1 into the Keap1-Cul3-E3 ligase complex, enhancing complex stability and enzymatic activity. This leads to increased Nrf2 protein degradation and reduced Nrf2 target gene expression, suppressing the antioxidant response. In vitro deubiquitylation assay, co-immunoprecipitation, siRNA knockdown, Western blot, reporter gene assays Molecular cell High 23727018
2014 USP15 stabilizes the E3 ubiquitin ligase MDM2 through deubiquitination. USP15-stabilized MDM2 negatively regulates T cell activation by targeting NFATc2 for degradation. USP15 deficiency promotes T cell activation and enhances antitumor T cell responses. USP15 also stabilizes MDM2 in cancer cells to regulate p53 function. Co-IP, in vitro deubiquitylation assay, USP15 knockout mice, T cell activation assays, tumor challenge models Nature immunology High 24777531
2014 USP15 opposes Parkin-mediated mitophagy by counteracting Parkin-mediated mitochondrial ubiquitination. USP15 does not affect Parkin ubiquitination status or Parkin translocation to mitochondria, but specifically counteracts ubiquitination of outer mitochondrial membrane proteins. A catalytically inactive version of USP15 does not oppose mitophagy. Knockdown of USP15 rescues the mitophagy defect in PD patient fibroblasts with PARK2 mutations. siRNA knockdown, catalytically inactive mutant, mitophagy assays, patient fibroblasts, Drosophila parkin RNAi rescue Human molecular genetics High 24852371
2014 USP15 deubiquitylates TRIM25, preventing LUBAC-dependent K48-linked polyubiquitylation and proteasomal degradation of TRIM25. USP15 was identified as an interaction partner of TRIM25 by protein purification and mass spectrometry. Wild-type but not catalytically inactive USP15 reduces K48-linked ubiquitylation of TRIM25, stabilizes it, and enhances TRIM25- and RIG-I-dependent type I IFN production. Affinity purification, mass spectrometry, Co-IP, siRNA knockdown, overexpression of catalytically inactive mutant, IFN reporter assays, viral replication assays Science signaling High 24399297
2014 USP15 identifies and deubiquitylates monoubiquitinated histone H2B (ubH2B). In the nucleus, USP15 indirectly associates with the ubH2B E3 ligase RNF20/RNF40, and directly associates with SART3 (TIP110/p110). SART3 acts as a histone chaperone that enhances USP15 binding to ubH2B and facilitates deubiquitination of free ubH2B histones. Affinity purification with nonhydrolyzable ubH2B mimic, Co-IP, in vitro deubiquitination assay with SART3 The Journal of biological chemistry High 24526689
2014 USP15 deubiquitylates ALK3/BMPR1A (BMP type I receptor) by interacting with SMAD6, an inhibitory SMAD that recruits E3 ubiquitin ligases to ALK3. USP15 reduces K48-linked polyubiquitylation of ALK3, enhances BMP-induced SMAD1 phosphorylation, and promotes BMP target gene transcription. Loss of USP15 inhibits BMP-induced osteoblast differentiation in myoblasts and modulates BMP signaling in Xenopus embryos. Co-IP, RNAi depletion, ubiquitination assay, SMAD1 phosphorylation assay, BMP target gene reporter, Xenopus embryo experiments Open biology High 24850914
2015 USP15 deubiquitinates SMURF2 at Lys734, a residue required for SMURF2 catalytic activity. This results in enhanced TβR-I stability and downstream TGF-β pathway activation. Proteomic analysis identified SMURF2 as a USP15 target in addition to TβR-I. Proteomic mass spectrometry, Co-IP, in vitro deubiquitination assay, mutational analysis of Lys734 Scientific reports Medium 26435193
2015 USP4, USP15, and USP11 arose by whole genome and small-scale duplications in vertebrate evolution. Viability of mice is contingent on a functional copy of either USP4 or USP15 (genetic redundancy established by cross-breeding mice with inactivating mutations in both genes). Phylogenetic and syntenic reconstruction, genetic crosses of Usp4/Usp15 mutant mice BMC evolutionary biology Medium 26503449
2016 USP15 co-expressed with and functionally acts together with the E3 ubiquitin ligase TRIM25 to positively regulate type I interferon responses. The USP15L749R mutation in mice dampens type I interferon responses in brain and hematopoietic cells, protecting against experimental cerebral malaria and EAE neuroinflammation. ENU-mutagenesis mouse model, immunophenotyping, RNA sequencing, in situ neuroinflammation models Nature immunology High 27721430
2016 TGF-β promotes translation of USP15 through activation of mTOR via the PI3K/AKT pathway. Upregulated USP15 then binds to and stabilizes p53 through deubiquitination in U2OS and HEK293 cells. Co-IP, ubiquitination assay, PI3K/AKT/mTOR pathway inhibitors, Western blot Oncogene Medium 27893708
2016 Structural basis for SART3 recruitment of USP15: crystal structures of SART3 alone and in complex with USP15 DUSP-UBL domain at 2.0 and 3.0 Å respectively reveal SART3 contains 12 HAT repeats in two subdomains, dimerizes through HAT-C concave surface, and binds USP15 in a novel bipartite mode on the HAT-C convex surface. USP15 binds SART3 ~20-fold more tightly than USP4. X-ray crystallography, isothermal titration calorimetry, mutagenesis The Journal of biological chemistry High 27255711
2017 USP15, together with its substrate-targeting factor SART3, deubiquitinates PRP31 (a U4 snRNP component). PRP31 is modified with K63-linked ubiquitin chains by the PRP19 complex. USP15-SART3 forms a complex with USP4, and this ternary complex also deubiquitinates PRP3. The ubiquitination/deubiquitination status of PRP31 regulates its interaction with U5 snRNP component PRP8, required for efficient splicing of chromosome segregation-related genes. Co-IP, in vitro deubiquitination assay, ubiquitin linkage-specific analysis, splicing assays Nucleic acids research High 28088760
2017 The long isoform of USP15 (containing exon 7-encoded serine-rich stretch) preferentially recognizes and deubiquitylates mysterin/RNF213 (a large ubiquitin ligase associated with moyamoya disease), while the short isoform does not. Exon 7 skipping alters substrate specificity of USP15 isoforms, with only partially overlapping interactomes. Isoform-specific pulldown, deubiquitylation assay, mass spectrometry interactome comparison Scientific reports Medium 28276505
2017 USP15 interacts with and deubiquitinates SLIM1 (skeletal muscle LIM protein 1), increasing SLIM1 protein levels. Cardiac-specific overexpression of USP15 in transgenic mice induces cardiac remodeling with elevated heart weight/body weight ratios and upregulation of fetal gene markers, associated with increased endogenous SLIM1 levels. Cell-free binding, co-immunoprecipitation, in vitro deubiquitination, transgenic mouse model Biochemical and biophysical research communications Medium 21219870
2018 Crystal structure of the USP15 catalytic core domain reveals a canonical USP fold (finger, palm, thumb regions) but with a misaligned catalytic triad — catalytic cysteine ~10 Å from catalytic histidine, an inactive configuration. Active-site loops are flexible, creating a largely open ubiquitin tail-binding channel. USP15 displays lower monoubiquitin affinity than paralog USP4. Mitoxantrone weakly inhibits USP15 and binds the S1' site. X-ray crystallography, isothermal titration calorimetry, enzyme inhibition assay The Journal of biological chemistry High 30228188
2018 USP15 deubiquitylates BARD1 BRCT domain upon recruitment to DNA double-strand breaks (DSBs) by MDC1. Recruitment requires the FHA domain of MDC1 and phosphorylated Ser678 of USP15. USP15-mediated BARD1 deubiquitylation promotes BARD1-HP1γ interaction, retaining BRCA1/BARD1 at DSBs and promoting homologous recombination (HR). USP15 knockout mice exhibit genomic instability. Co-IP, phospho-mutant analysis, BARD1 deubiquitylation assay, HR assay, USP15 KO mice Nature communications High 30874560
2018 USP15 deubiquitylates topoisomerase IIα (TOP2A), and USP15 depletion leads to decreased TOP2A accumulation, formation of anaphase chromosome bridges, and micronuclei indicating genome instability. Both USP15 isoforms move from cytoplasm to nucleus at prophase; isoform-1 is phosphorylated on S229 at mitotic entry and an S229D phospho-mimetic cannot rescue TOP2A accumulation or the micronuclei phenotype, demonstrating isoform-specific phospho-regulation. siRNA depletion, phospho-mutant rescue, live-cell imaging, cell cycle analysis, ubiquitination assay Oncogene High 29429988
2018 HPV16 E6 oncoprotein forms a complex with TRIM25 and USP15 in human cells. In the presence of E6, K48-linked ubiquitination of TRIM25 is markedly increased and TRIM25 degradation is enhanced, and E6 inhibits TRIM25-mediated K63-linked ubiquitination of RIG-I and its CARD-dependent interaction with MAVS. Co-IP, ubiquitination assay, CRISPR-Cas9 knockout in keratinocytes, IFN reporter assay Journal of virology Medium 29263274
2019 Ubiquitin variants (UbVs) targeting either the USP15 catalytic domain or each of three adaptor domains (including the DUSP domain) inhibit USP15 catalytic activity. A linear dimer UbV targeting both DUSP and catalytic domains shows enhanced specificity. Crystal structures show three distinct UbVs bind the catalytic domain and lock the active site in a closed, inactive conformation; one UbV forms an unusual strand-swapped dimer binding two DUSP domains simultaneously. In cells, UbVs inhibit deubiquitination of SMURF2 and TRIM25 substrates. Protein engineering (UbVs), X-ray crystallography, cell-based deubiquitination assays, TGF-β pathway assays Structure (London, England : 1993) High 30713027
2019 The ER-located E3 ligase Hrd1 interacts with USP15 and ubiquitinates it. Unlike classical Hrd1 substrates, USP15 is not degraded but loses its deubiquitinating activity toward IκBα, resulting in excessive NF-κB activation. This represents a non-canonical Hrd1 function linking ER-plasma membrane signaling during bacterial TLR4 responses. E3 ligase screen (>280 ligases), Co-IP, ubiquitination assay, IκBα deubiquitination assay, macrophage-specific Hrd1 KO mice, LPS-sepsis model Nature microbiology High 31477895
2019 USP15 phosphorylation at Thr149 and Thr219 (identified by nuclear-cytoplasmic fractionation and mass spectrometry) occurs in the cytoplasm. The phosphorylation status alters interaction with SART3, leading to nuclear localization and deubiquitinating activity toward spliceosomal substrate PRP31. Treatment with CDK inhibitor purvalanol A induces nuclear translocation of USP15. Subcellular fractionation, mass spectrometry, phospho-mutant analysis, Co-IP, in vitro deubiquitination assay Journal of molecular biology Medium 31330151
2019 USP15 deubiquitylates and stabilizes HECTD1 E3 ubiquitin ligase in glioblastoma cells. USP15 expression attenuates canonical WNT pathway activity in a manner dependent on HECTD1; depletion of USP15 reduces HECTD1 protein levels. USP15 interaction with HECTD1 was identified by mass spectrometry. Mass spectrometry protein network analysis, Co-IP, ubiquitination assay, WNT reporter assay, soft agar colony formation Oncotarget Medium 29299163
2020 USP15 deubiquitylates and inactivates TET2 by removing K1299-linked monoubiquitin, which normally promotes TET2 activity. Deletion of Usp15 in melanoma cells stimulates chemokine expression and tumor-infiltrating lymphocyte accumulation in a TET2-dependent manner, leading to increased immunotherapy response. Co-IP, in vitro deubiquitylation assay, gene expression profiling, Usp15 KO melanoma model, immunotherapy challenge Science advances High 32948596
2020 USP15 potentiates NF-κB activation upon TNFα or IL-1β stimulation by stabilizing TAB2 and TAB3 through different mechanisms: (1) catalytic-activity-dependent deubiquitination of K48-linked TAB2 ubiquitin chains; (2) catalytic-activity-independent inhibition of lysosome-mediated TAB2 degradation; (3) catalytic-activity-independent inhibition of NBR1-mediated selective autophagic TAB3 degradation. Co-IP, ubiquitination assay, overexpression/knockdown of USP15, catalytically inactive mutant, NF-κB reporter, lysosome/autophagy inhibitor experiments The FEBS journal Medium 31903660
2020 UBE2S recruits USP15 to TBK1 to remove K63-linked polyubiquitin chains from TBK1, thereby inhibiting TBK1 activation and type I IFN production. UBE2S-mediated inhibition of IFN is independent of its E2/E3 enzymatic activity but requires USP15 recruitment. Co-IP, ubiquitination assay (K63 linkage-specific), USP15 knockdown, viral replication assays in vitro and in vivo Cell reports Medium 32814047
2020 TIFAB regulates USP15 ubiquitin hydrolase activity (acts as an effector/activator of USP15). Expression of TIFAB in hematopoietic stem/progenitor cells permits USP15 signaling to substrates MDM2 and KEAP1, mitigating p53 expression. TIFAB-deficient HSPCs exhibit compromised USP15 signaling and are sensitized to hematopoietic stress by derepression of p53. Proteomics, co-IP, USP15 activity assays, genetic knockdown/overexpression in HSPCs, MLL-AF9 leukemia model Cell reports Medium 32101751
2020 USP15 interacts with and stabilizes FUS (fused in sarcoma), a known DNA repair factor, in leukemia cells. USP15 is essential for HSC maintenance in vitro and in vivo (transplantation and Usp15 KO mice), and its depletion in leukemia cells impairs expansion and increases genotoxic stress. Co-IP, shRNA in vivo screen, Usp15 KO mice, transplantation assays, genotoxic stress assays Cell reports Medium 33378683
2020 USP15 deubiquitinates CARD9, constitutively associating with it and removing TRIM62-deposited ubiquitin marks. USP15 knockdown and knockout specifically enhance CARD9-dependent C-type lectin receptor (CLR) signaling in mouse and human immune cells. Co-IP, ubiquitination assay, siRNA knockdown, CRISPR KO, CLR signaling assays ImmunoHorizons Medium 33093067
2021 USP15 deubiquitinates and stabilizes glutamine synthetase (GS) and CRL4CRBN neosubstrates IKZF1, IKZF3, CK1-α, RNF166, GSPT1, and BRD4 by antagonizing CRL4CRBN-mediated ubiquitylation. USP15 is highly expressed in IMiD-resistant myeloma cells and depletion sensitizes these cells to lenalidomide. Ubiquitination assays, Western blot for substrate stability, siRNA depletion, lenalidomide sensitivity assays Proceedings of the National Academy of Sciences of the United States of America Medium 34583995
2021 USP15 deubiquitinates and stabilizes p53-R175H gain-of-function mutant through a lysosome-mediated pathway. USP15 is established as a selective upstream regulator of p53-R175H stability distinct from wild-type p53 regulation. Chemical genetic approach (MCB-613), siRNA knockdown, CRISPR, ubiquitination assay, lysosome pathway assays Nature communications Medium 29593334
2021 USP15 deubiquitinates TUT1 (terminal uridylyl transferase 1) and causes redistribution of TUT1 from nucleolus to nucleoplasm, stabilizing U6 snRNA. Usp15 knockout mice show impaired motor ability and unconventional cerebellar formation. Inhibition of the USP15-TUT1 cascade triggers mild chronic ER stress. Co-IP, ubiquitination assay, immunofluorescence localization, Usp15 KO mice, U6 snRNA stability assay, ER stress markers Molecular and cellular biology Medium 32839293
2021 USP15 deubiquitinates and stabilizes ERα (estrogen receptor alpha) by removing K48-linked ubiquitin, promoting ERα+ breast cancer cell proliferation. USP15 knockdown reduces ERα protein levels and enhances tamoxifen antitumor activity. Co-IP, ubiquitination assay (K48 linkage-specific), siRNA knockdown, in vitro and in vivo proliferation assays Cell death & disease Medium 33771975
2021 USP15 deubiquitinates BECN1 (Beclin-1), interacting with BECN1 but not TRAF6, thereby attenuating TRAF6-BECN1 axis-driven autophagy induction. USP15-knockout lung cancer cells show increased cancer migration and invasion with enhanced autophagy in response to TLR4 stimulation. Co-IP, deubiquitination assay, CRISPR-Cas9 KO cell lines, migration/invasion assays, autophagy assays Cell death & disease Medium 35422093
2022 USP15 interacts with and promotes cGAS activation through deubiquitylation of cGAS and promotion of cGAS dimerization and liquid condensation via the USP15 intrinsic disordered region (IDR). In the absence of DNA, USP15 drives cGAS liquid condensation preparing cGAS for rapid DNA response. Co-IP, in vitro deubiquitylation assay, liquid-liquid phase separation assay, IDR deletion mutant analysis, cGAS activation assays Nucleic acids research Medium 36243958
2022 Crystal structure of USP15 D1D2 catalytic domain in a catalytically competent conformation shows the active site can switch between active and inactive states independently of mitoxantrone binding. Mitoxantrone contributes to crystal packing by forming a stack of 12 molecules at the S1' site rather than inducing catalytic triad misalignment. X-ray crystallography (multiple crystal structures), comparative structural analysis Journal of structural biology High 35605756
2023 USP15 interacts with and deubiquitinates PARP1 to promote its stability, stimulating DNA repair, genomic stability, and TNBC cell proliferation. ER inhibits USP15 expression at the promoter level, PR suppresses USP15 deubiquitinase activity, and HER2 abrogates the PARP1-USP15 interaction — explaining elevated PARP1 in TNBC where all three receptors are absent. Co-IP, in vitro deubiquitination assay, ChIP, USP15 activity assay in presence of PR, Co-IP in presence of HER2, TNBC cell proliferation/DNA repair assays Nature cancer High 37012401
2023 The CRL3gigaxonin E3 ligase and USP15 form an opposing regulatory pathway governing destruction of neurofilament proteins NEFL and INA. USP15 antagonizes CRL3GIG-mediated ubiquitylation of NEFL and INA. A specific degron (NEFLL12) in neurofilament proteins is required for CRL3GIG binding; mutations in the GIG Kelch domain (L309R, R545C, C570Y) disrupt NEFL/INA binding causing accumulation of NF proteins as in giant axonal neuropathy. Ubiquitination assays, Co-IP, degron mapping, mutant GIG binding assays, proteasome degradation assays Proceedings of the National Academy of Sciences of the United States of America Medium 37903270
2023 USP15 deubiquitinates SMYD3, stabilizing it and enabling H3K4me3-mediated transcriptional activation of CCL2. This promotes MDSC recruitment to tumors and immune evasion in colorectal cancer. USP15 inhibition improves anti-PD-1 efficacy in colorectal cancer models. Co-IP, deubiquitination assay, H3K4me3 ChIP, MDSC recruitment assays, orthotopic/metastatic colon tumor models, PD-1 blockade combination Cancer immunology research Medium 40323348
2023 USP15 interacts with and deubiquitinates YAP1, inhibiting K48-linked ubiquitination of YAP1 and stabilizing it. USP15 also binds TAZ after hypoxia treatment. USP15 promotes PASMC proliferation and migration in a YAP1/TAZ-dependent manner, contributing to pulmonary vascular remodeling. Co-IP, ubiquitination assay (K48 linkage), USP15 knockdown (AAV-mediated in vivo), YAP1/TAZ rescue experiments, SuHx and MCT mouse models Experimental & molecular medicine Medium 36635430
2024 USP15 deubiquitinates HKDC1 (a glycolytic regulator), inhibiting its ubiquitination-mediated degradation and thereby regulating glucose metabolism and glycolytic activity in gastric cancer cells. Co-IP, ubiquitination assay, USP15 knockdown/overexpression, glycolytic activity assays, xenograft models Journal of experimental & clinical cancer research : CR Medium 39164728
2024 TRIM21 E3 ligase and USP15 antagonistically regulate ACSL4 protein stability in gastrointestinal stromal tumors (GISTs); TRIM21-mediated ubiquitination decreases ACSL4 while USP15 opposes this, and the balance determines imatinib sensitivity. Co-IP, ubiquitination assay, Western blot for ACSL4 stability, shRNA, xenograft model, GIST patient clinical data British journal of cancer Medium 38182686

Source papers

Stage 0 corpus · 100 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2012 USP15 stabilizes TGF-β receptor I and promotes oncogenesis through the activation of TGF-β signaling in glioblastoma. Nature medicine 347 22344298
2014 The deubiquitinase USP15 antagonizes Parkin-mediated mitochondrial ubiquitination and mitophagy. Human molecular genetics 265 24852371
2014 USP15 stabilizes MDM2 to mediate cancer-cell survival and inhibit antitumor T cell responses. Nature immunology 210 24777531
2011 USP15 is a deubiquitylating enzyme for receptor-activated SMADs. Nature cell biology 165 21947082
2014 The ubiquitin-specific protease USP15 promotes RIG-I-mediated antiviral signaling by deubiquitylating TRIM25. Science signaling 139 24399297
2018 The Human Papillomavirus E6 Oncoprotein Targets USP15 and TRIM25 To Suppress RIG-I-Mediated Innate Immune Signaling. Journal of virology 126 29263274
2005 The zinc finger of the CSN-associated deubiquitinating enzyme USP15 is essential to rescue the E3 ligase Rbx1. Current biology : CB 126 16005295
2013 USP15 negatively regulates Nrf2 through deubiquitination of Keap1. Molecular cell 103 23727018
2009 The COP9 signalosome mediates beta-catenin degradation by deneddylation and blocks adenomatous polyposis coli destruction via USP15. Journal of molecular biology 94 19576224
2019 The deubiquitylating enzyme USP15 regulates homologous recombination repair and cancer cell response to PARP inhibitors. Nature communications 93 30874560
2014 The U4/U6 recycling factor SART3 has histone chaperone activity and associates with USP15 to regulate H2B deubiquitination. The Journal of biological chemistry 83 24526689
2016 USP15 regulates type I interferon response and is required for pathogenesis of neuroinflammation. Nature immunology 75 27721430
2016 TGF-β upregulates the translation of USP15 via the PI3K/AKT pathway to promote p53 stability. Oncogene 75 27893708
2018 USP15-dependent lysosomal pathway controls p53-R175H turnover in ovarian cancer cells. Nature communications 71 29593334
2022 USP15 negatively regulates lung cancer progression through the TRAF6-BECN1 signaling axis for autophagy induction. Cell death & disease 70 35422093
2009 The ubiquitin-specific peptidase USP15 regulates human papillomavirus type 16 E6 protein stability. Journal of virology 67 19553310
2020 USP15 potentiates NF-κB activation by differentially stabilizing TAB2 and TAB3. The FEBS journal 60 31903660
1999 Identification, functional characterization, and chromosomal localization of USP15, a novel human ubiquitin-specific protease related to the UNP oncoprotein, and a systematic nomenclature for human ubiquitin-specific proteases. Genomics 59 10444327
2019 IL1R2 Blockade Suppresses Breast Tumorigenesis and Progression by Impairing USP15-Dependent BMI1 Stability. Advanced science (Weinheim, Baden-Wurttemberg, Germany) 57 31921558
2023 Loss of the receptors ER, PR and HER2 promotes USP15-dependent stabilization of PARP1 in triple-negative breast cancer. Nature cancer 55 37012401
2015 Evolution of the highly networked deubiquitinating enzymes USP4, USP15, and USP11. BMC evolutionary biology 54 26503449
2009 USP15 plays an essential role for caspase-3 activation during Paclitaxel-induced apoptosis. Biochemical and biophysical research communications 52 19665996
2017 The Regulations of Deubiquitinase USP15 and Its Pathophysiological Mechanisms in Diseases. International journal of molecular sciences 50 28245560
2019 Structural and Functional Characterization of Ubiquitin Variant Inhibitors of USP15. Structure (London, England : 1993) 48 30713027
2019 ER-localized Hrd1 ubiquitinates and inactivates Usp15 to promote TLR4-induced inflammation during bacterial infection. Nature microbiology 48 31477895
2018 The structure of the deubiquitinase USP15 reveals a misaligned catalytic triad and an open ubiquitin-binding channel. The Journal of biological chemistry 48 30228188
2020 Ubiquitin-Conjugating Enzyme 2S Enhances Viral Replication by Inhibiting Type I IFN Production through Recruiting USP15 to Deubiquitinate TBK1. Cell reports 46 32814047
2015 USP15 regulates SMURF2 kinetics through C-lobe mediated deubiquitination. Scientific reports 46 26435193
2015 T Cell Intrinsic USP15 Deficiency Promotes Excessive IFN-γ Production and an Immunosuppressive Tumor Microenvironment in MCA-Induced Fibrosarcoma. Cell reports 46 26686633
2020 USP15 suppresses tumor immunity via deubiquitylation and inactivation of TET2. Science advances 45 32948596
2014 USP15 targets ALK3/BMPR1A for deubiquitylation to enhance bone morphogenetic protein signalling. Open biology 44 24850914
2021 USP15 antagonizes CRL4CRBN-mediated ubiquitylation of glutamine synthetase and neosubstrates. Proceedings of the National Academy of Sciences of the United States of America 43 34583995
2012 Direct and indirect control of mitogen-activated protein kinase pathway-associated components, BRAP/IMP E3 ubiquitin ligase and CRAF/RAF1 kinase, by the deubiquitylating enzyme USP15. The Journal of biological chemistry 42 23105109
2013 The deubiquitylase USP15 stabilizes newly synthesized REST and rescues its expression at mitotic exit. Cell cycle (Georgetown, Tex.) 41 23708518
2021 The deubiquitinating enzyme USP15 stabilizes ERα and promotes breast cancer progression. Cell death & disease 39 33771975
2021 USP15: a review of its implication in immune and inflammatory processes and tumor progression. Genes and immunity 39 33824497
2021 The Multifaceted Roles of USP15 in Signal Transduction. International journal of molecular sciences 38 33946990
2020 De-regulated STAT5A/miR-202-5p/USP15/Caspase-6 regulatory axis suppresses CML cell apoptosis and contributes to Imatinib resistance. Journal of experimental & clinical cancer research : CR 38 31952546
2018 USP15 inhibits multiple myeloma cell apoptosis through activating a feedback loop with the transcription factor NF-κBp65. Experimental & molecular medicine 35 30459344
2020 TIFAB Regulates USP15-Mediated p53 Signaling during Stressed and Malignant Hematopoiesis. Cell reports 34 32101751
2018 The deubiquitylase USP15 regulates topoisomerase II alpha to maintain genome integrity. Oncogene 34 29429988
2022 USP15 in Cancer and Other Diseases: From Diverse Functionsto Therapeutic Targets. Biomedicines 33 35203682
2021 USP15 Enhances the Proliferation, Migration, and Collagen Deposition of Hypertrophic Scar-Derived Fibroblasts by Deubiquitinating TGF-βR1 In Vitro. Plastic and reconstructive surgery 31 34546211
2017 USP15 regulates dynamic protein-protein interactions of the spliceosome through deubiquitination of PRP31. Nucleic acids research 31 28088760
2017 Ubiquitin Specific Peptidase 15 (USP15) suppresses glioblastoma cell growth via stabilization of HECTD1 E3 ligase attenuating WNT pathway activity. Oncotarget 30 29299163
2023 PLOD2 promotes colorectal cancer progression by stabilizing USP15 to activate the AKT/mTOR signaling pathway. Cancer science 29 37227305
2021 The deubiquitinase USP15 modulates cellular redox and is a therapeutic target in acute myeloid leukemia. Leukemia 29 34465865
2022 GINS1 promotes the proliferation and migration of glioma cells through USP15-mediated deubiquitination of TOP2A. iScience 28 36065190
2017 Deubiquitylation of hepatitis B virus X protein (HBx) by ubiquitin-specific peptidase 15 (USP15) increases HBx stability and its transactivation activity. Scientific reports 25 28074857
2021 Platelet-Rich Plasma-Derived Exosomal USP15 Promotes Cutaneous Wound Healing via Deubiquitinating EIF4A1. Oxidative medicine and cellular longevity 24 34422211
2019 USP15 Participates in Hepatitis C Virus Propagation through Regulation of Viral RNA Translation and Lipid Droplet Formation. Journal of virology 23 30626683
2019 Phosphorylation of USP15 and USP4 Regulates Localization and Spliceosomal Deubiquitination. Journal of molecular biology 23 31330151
2016 Structural Basis of the Recruitment of Ubiquitin-specific Protease USP15 by Spliceosome Recycling Factor SART3. The Journal of biological chemistry 23 27255711
2021 Positive feedback regulation between USP15 and ERK2 inhibits osteoarthritis progression through TGF-β/SMAD2 signaling. Arthritis research & therapy 22 33726807
2011 Structure of the USP15 N-terminal domains: a β-hairpin mediates close association between the DUSP and UBL domains. Biochemistry 22 21848306
2023 USP15 promotes pulmonary vascular remodeling in pulmonary hypertension in a YAP1/TAZ-dependent manner. Experimental & molecular medicine 21 36635430
2022 USP15 and USP4 facilitate lung cancer cell proliferation by regulating the alternative splicing of SRSF1. Cell death discovery 21 35027535
2019 COP9 Signalosome Interaction with UspA/Usp15 Deubiquitinase Controls VeA-Mediated Fungal Multicellular Development. Biomolecules 21 31216760
2014 Regulation of ubiquitin-proteasome system-mediated Tip110 protein degradation by USP15. The international journal of biochemistry & cell biology 21 24984263
2022 USP15 promotes cGAS activation through deubiquitylation and liquid condensation. Nucleic acids research 20 36243958
2020 USP15 promotes the apoptosis of degenerative nucleus pulposus cells by suppressing the PI3K/AKT signalling pathway. Journal of cellular and molecular medicine 20 33135363
2020 USP15 Deubiquitinase Safeguards Hematopoiesis and Genome Integrity in Hematopoietic Stem Cells and Leukemia Cells. Cell reports 20 33378683
2023 USP15 Represses Hepatocellular Carcinoma Progression by Regulation of Pathways of Cell Proliferation and Cell Migration: A System Biology Analysis. Cancers 19 36900163
2020 Inhibition of USP15 Prevent Glutamate-Induced Oxidative Damage by Activating Nrf2/HO-1 Signaling Pathway in HT22 Cells. Cellular and molecular neurobiology 19 31933062
2023 Ablation of the deubiquitinase USP15 ameliorates nonalcoholic fatty liver disease and nonalcoholic steatohepatitis. Experimental & molecular medicine 18 37394587
2021 USP15 participates in DBP-induced testicular oxidative stress injury through regulating the Keap1/Nrf2 signaling pathway. The Science of the total environment 18 34088152
2011 Transgenic overexpression of USP15 in the heart induces cardiac remodeling in mice. Biochemical and biophysical research communications 17 21219870
2003 Isolation and characterization of the mouse ubiquitin-specific protease Usp15. Mammalian genome : official journal of the International Mammalian Genome Society 17 12532266
2024 PRDM1 promotes the ferroptosis and immune escape of thyroid cancer by regulating USP15-mediated SELENBP1 deubiquitination. Journal of endocrinological investigation 16 39014173
2023 ACVRL1 drives resistance to multitarget tyrosine kinase inhibitors in colorectal cancer by promoting USP15-mediated GPX2 stabilization. BMC medicine 16 37743483
2022 USP15 regulates p66Shc stability associated with Drp1 activation in liver ischemia/reperfusion. Cell death & disease 16 36163170
2020 USP15 Enhances Re-epithelialization Through Deubiquitinating EIF4A1 During Cutaneous Wound Repair. Frontiers in cell and developmental biology 16 32671073
2017 USP15 attenuates IGF-I signaling by antagonizing Nedd4-induced IRS-2 ubiquitination. Biochemical and biophysical research communications 16 28126338
2016 Function of ubiquitin (Ub) specific protease 15 (USP15) in HIV-1 replication and viral protein degradation. Virus research 16 27460547
2013 Zebrafish transforming growth factor-β-stimulated clone 22 domain 3 (TSC22D3) plays critical roles in Bmp-dependent dorsoventral patterning via two deubiquitylating enzymes Usp15 and Otud4. Biochimica et biophysica acta 16 23665588
2024 TRIM21/USP15 balances ACSL4 stability and the imatinib resistance of gastrointestinal stromal tumors. British journal of cancer 15 38182686
2020 USP15 Deubiquitinates TUT1 Associated with RNA Metabolism and Maintains Cerebellar Homeostasis. Molecular and cellular biology 15 32839293
2017 Alternative exon skipping biases substrate preference of the deubiquitylase USP15 for mysterin/RNF213, the moyamoya disease susceptibility factor. Scientific reports 14 28276505
2024 The deubiquitinase USP15 drives malignant progression of gastric cancer through glucose metabolism remodeling. Journal of experimental & clinical cancer research : CR 13 39164728
2023 FUNDC1/USP15/Drp1 ameliorated TNF-α-induced pulmonary artery endothelial cell proliferation by regulating mitochondrial dynamics. Cellular signalling 12 37871666
2023 miR-196a provides antioxidative neuroprotection via USP15/Nrf2 regulation in Huntington's disease. Free radical biology & medicine 12 37907121
2021 Inhibition of USP15 ameliorates high-glucose-induced oxidative stress and inflammatory injury in podocytes through regulation of the Keap1/Nrf2 signaling. Environmental toxicology 11 34931430
2020 USP15 Deubiquitinates CARD9 to Downregulate C-Type Lectin Receptor-Mediated Signaling. ImmunoHorizons 10 33093067
2024 K48-linked deubiquitination of VGLL4 by USP15 enhances the efficacy of tumor immunotherapy in triple-negative breast cancer. Cancer letters 9 38431034
2014 Expression of USP15, TβR-I and Smad7 in psoriasis. Journal of Huazhong University of Science and Technology. Medical sciences = Hua zhong ke ji da xue xue bao. Yi xue Ying De wen ban = Huazhong keji daxue xuebao. Yixue Yingdewen ban 9 24939309
2025 N6-methyladenosine-modification of USP15 regulates chemotherapy resistance by inhibiting LGALS3 ubiquitin-mediated degradation via AKT/mTOR signaling activation pathway in hepatocellular carcinoma. Cell death discovery 8 39794359
2023 The CRL3gigaxonin ubiquitin ligase-USP15 pathway governs the destruction of neurofilament proteins. Proceedings of the National Academy of Sciences of the United States of America 8 37903270
2022 Mitoxantrone stacking does not define the active or inactive state of USP15 catalytic domain. Journal of structural biology 8 35605756
2021 KIF15 upregulation promotes leiomyosarcoma cell growth via promoting USP15-mediated DEK deubiquitylation. Biochemical and biophysical research communications 8 34280614
2018 USP15 inhibits HPV16 E6 degradation and catalytically inactive USP15 has reduced inhibitory activity. Acta virologica 8 29895155
2025 USP15 Facilitates Colorectal Cancer Immune Evasion through SMYD3/CCL2-Dependent Myeloid-Derived Suppressor Cell Recruitment. Cancer immunology research 7 40323348
2024 USP15, activated by TFAP4 transcriptionally, stabilizes SHC1 via deubiquitination and deteriorates renal cell carcinoma. Cancer science 7 38847328
2023 LncRNA ZNNT1 induces p53 degradation by interfering with the interaction between p53 and the SART3-USP15 complex. PNAS nexus 7 37448957
2022 Tip110/SART3-Mediated Regulation of NF-κB Activity by Targeting IκBα Stability Through USP15. Frontiers in oncology 7 35530338
2022 SRA Suppresses Antiviral Innate Immune Response in Macrophages by Limiting TBK1 K63 Ubiquitination via Deubiquitinase USP15. Microbiology spectrum 7 36342281
2023 Hypoxia-induced paclitaxel resistance in cervical cancer modulated by miR-100 targeting of USP15. Gynecologic oncology reports 6 36714373
2025 Narirutin downregulates USP15 to inhibit the NLRP3 ubiquitination in HDM-induced allergic rhinitis. Phytomedicine : international journal of phytotherapy and phytopharmacology 5 40517692
2024 USP15 promotes the progression of papillary thyroid cancer by regulating HMGB1 stability through its deubiquitination. Journal of Cancer 5 38577597
2023 miR-26a exerts broad-spectrum antiviral effects via the enhancement of RIG-I-mediated type I interferon response by targeting USP15. Microbiology spectrum 5 38019020
2022 Alleviative effect of microRNA-497 on diabetic neuropathic pain in rats in relation to decreased USP15. Cell biology and toxicology 5 35478295

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