Affinage

RBFOX2

RNA binding protein fox-1 homolog 2 · UniProt O43251

Length
390 aa
Mass
41.4 kDa
Annotated
2026-06-10
90 papers in source corpus 50 papers cited in narrative 50 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 7/7 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

RBFOX2 is a sequence-specific RNA-binding protein that governs alternative splicing programs across development, tissue homeostasis, and disease by recognizing the hexameric element UGCAUG (and related GCAUG motifs) in pre-mRNAs (PMID:16537540, PMID:18794351). Its regulatory outcome is positional: binding downstream of an alternative exon activates inclusion while upstream binding represses it (PMID:18794351), and mechanistically it both recruits U1 snRNP to weak 5' splice sites through a direct C-terminal-domain interaction with U1C (PMID:22083953) and blocks prespliceosome assembly at two distinct steps by occluding U2AF65/SF1-dependent complex formation (PMID:17101796, PMID:18573872). Beyond cassette-exon splicing, RBFOX2 controls AS-coupled NMD of RNA-binding protein transcripts to tune a broader RBP network (PMID:24637117), directs alternative polyadenylation (PMID:34731606), and acts in the nucleus on chromatin: it binds PRC2 in a nascent-RNA-dependent manner to enforce silencing at bivalent promoters (PMID:27211866) and recognizes m6A on chromatin-associated RNAs to recruit an RBM15/YTHDC1/PRC2 axis for locus-selective gene silencing (PMID:37640841). RBFOX2 cooperates and competes with other splicing regulators including hnRNPC, hnRNPM and SRSF1 in distinct binding configurations (PMID:34244793), MBNL1 and QKI (PMID:24048253, PMID:40207498), and its activity is constrained by autoregulatory and trans-acting production of dominant-negative isoforms (PMID:16449636, PMID:21747913, PMID:27239029). Isoform-specific nuclear localization signals partition RBFOX2 between nuclear splicing functions and a cytoplasmic stress-granule role regulating RB1 mRNA (PMID:27859055, PMID:28894257, PMID:31028247). Its abundance is set post-translationally by competing ubiquitin pathways: FBXO7/PRMT5-dependent K63 ubiquitination stabilizes it (PMID:37822160) while it is a substrate of the RhoBTB1/CUL3 degradation pathway (PMID:41926228). Through these activities RBFOX2 directs cerebellar and Purkinje-cell pacemaking via sodium-channel splicing (PMID:22357600), myoblast fusion and satellite-cell activation (PMID:25087874, PMID:41418788), cardiac development and stress responses (PMID:25753418, PMID:30867288, PMID:35137168), pancreatic β-cell insulin secretion (PMID:38007492), HSC proteostasis (PMID:41337583), vascular endothelial and smooth-muscle function (PMID:29293084, PMID:41926228), liver cholesterol homeostasis (PMID:36536133), and suppression of metastasis in pancreatic and other cancers (PMID:36949200, PMID:38114498).

Mechanistic history

Synthesis pass · year-by-year structured walk · 18 steps
  1. 2006 High

    Established the core molecular activity—that RBFOX2 recognizes a defined RNA element and acts as a physiological splicing regulator—answering what RBFOX2 binds and what it does.

    Evidence SELEX defining UGCAUG specificity plus co-transfection splicing assays and siRNA knockdown on protein 4.1R and FGFR2 exons, with UGCAUG mutagenesis

    PMID:16449636 PMID:16537540

    Open questions at the time
    • Whether binding position alone dictates activate-vs-repress outcomes not yet generalized
    • C-terminal mechanism of activation not defined at this stage
  2. 2006 High

    Defined a repressive mechanism, showing RBFOX2 can occlude splice-site recognition rather than only activate inclusion.

    Evidence Overexpression in HeLa/neuronal cells with UGCAUG mutation analysis and U2AF65 binding assays on calcitonin/CGRP exon 4

    PMID:17101796

    Open questions at the time
    • Whether the same factor blocks more than one assembly step unresolved here
    • In vivo relevance of repression not tested
  3. 2008 Medium

    Resolved the repression mechanism into a two-step block of prespliceosome assembly and codified the positional rule that determines activation vs. repression.

    Evidence In vitro spliceosome assembly assays with UGCAUG mutants and gel-shift; genome-wide prediction with experimental RT-PCR validation

    PMID:18573872 PMID:18794351

    Open questions at the time
    • Positional rule rests partly on computational prediction
    • Quantitative determinants of position-dependence not fully mapped
  4. 2008 Medium

    Extended RBFOX2 beyond nuclear splicing by linking the ortholog to cytoplasmic polyadenylation, hinting at multifunctionality.

    Evidence Co-IP and tethered-function translation reporter in Xenopus oocytes (Rbm9 ortholog)

    PMID:18177378

    Open questions at the time
    • Single lab, Xenopus ortholog
    • Mammalian relevance of cytoplasmic polyadenylation role untested
  5. 2011 High

    Defined the activation mechanism at the molecular level—how RBFOX2 strengthens weak splice sites—and identified its cross-regulation by Rbfox3 through AS-NMD.

    Evidence Splice-site and CTD-deletion mutagenesis with U1C interaction/U1 snRNP recruitment assays; Rbfox3 isoform overexpression with NMD assays

    PMID:21747913 PMID:22083953

    Open questions at the time
    • Generality of U1C-mediated recruitment across targets not established
    • Rbfox3 cross-regulation tested in single study
  6. 2012 High

    Established an in vivo physiological requirement, showing RBFOX2 maintains neuronal excitability via sodium-channel splicing.

    Evidence CNS/Purkinje conditional knockout mice with RNA-seq, electrophysiology, and Nav1.6 immunoblot

    PMID:22357600

    Open questions at the time
    • Redundancy with Rbfox1 complicates Rbfox2-specific attribution
    • Direct binding to Scn8a not mapped at nucleotide resolution here
  7. 2014 High

    Expanded RBFOX2's role from individual exons to network-level control, defining AS-NMD of RBP transcripts and a myogenic splicing program required for myoblast fusion.

    Evidence iCLIP-seq plus RNA-seq in mouse ESCs and myoblasts, with NMD-isoform validation and Mef2d/Rock2 rescue of fusion defects

    PMID:24637117 PMID:25087874

    Open questions at the time
    • Hierarchy within the RBP regulatory network not fully ordered
    • Which targets drive fusion vs. are bystanders not exhaustively resolved
  8. 2015 High

    Established RBFOX2 as a stress-responsive splicing regulator in heart, where its loss recapitulates pressure-overload decompensation.

    Evidence Cardiac conditional knockout and TAC surgery with RNA-seq, RT-PCR, and echocardiography

    PMID:25753418

    Open questions at the time
    • Causal splicing targets driving decompensation not individually validated here
    • Mechanism linking pressure overload to RBFox2 downregulation unknown
  9. 2016 High

    Revealed a chromatin-level function distinct from splicing—RBFOX2 recruits PRC2 in a nascent-RNA-dependent manner to silence bivalent promoters.

    Evidence ChIP-seq, RBFox2–PRC2 co-IP, nascent RNA sequencing, and depletion across multiple cell types

    PMID:27211866

    Open questions at the time
    • How RBFox2 reads nascent RNA to dock PRC2 not structurally defined
    • Relationship between chromatin role and splicing role at same loci unclear
  10. 2016 Medium

    Defined isoform-encoded localization control and dominant-negative regulation, explaining how RBFOX2 partitions between nuclear and cytoplasmic functions and how disease isoforms inhibit wild-type activity.

    Evidence Isoform NLS mapping by subcellular fractionation/IF; DN-WT co-IP with calcium imaging in cardiomyocytes; HLHS truncation-mutant analysis with RIP and patient transcriptomes

    PMID:27239029 PMID:27485310 PMID:27859055

    Open questions at the time
    • DN isoform and HLHS findings from single studies
    • Quantitative threshold at which DN isoform overrides WT not defined
  11. 2017 Medium

    Identified a cytoplasmic stress-granule function in which RBFOX2 regulates RB1, decoupling a non-splicing role from its nuclear activity.

    Evidence IF co-localization with stress-granule markers and RIP-seq of cytoplasmic targets; calcium-channel splicing/electrophysiology in neurons and VSMCs

    PMID:28894257 PMID:28993448 PMID:29067356

    Open questions at the time
    • Mechanism of RBFOX2 recruitment into stress granules unknown
    • RB1 regulation tested in limited contexts
  12. 2018 High

    Established tissue-specific in vivo programs—vascular flow-responsive splicing in endothelium and craniofacial development via TGF-β-Tak1 splicing.

    Evidence Endothelial and neural-crest conditional knockouts with RNA-seq; Tak1 overexpression rescue and low-flow arterial model with immune-cell depletion

    PMID:29293084 PMID:31241461

    Open questions at the time
    • Direct splicing targets mediating each phenotype not all validated
    • Feedback loop architecture (TGF-β → RBFox2) mechanism partly inferred
  13. 2019 High

    Defined a feed-forward cardiac pathway (RBFox2 → miR-34a → Jph2) and confirmed stress-granule/cytoplasmic localization as a cancer-relevant axis.

    Evidence Cardiac KO with miR-34a mimic/antagomir rescue and Jph2 analysis; cancer-tissue IF with RB1 measurement and stress-granule dissolution assays

    PMID:30867288 PMID:31028247

    Open questions at the time
    • Mechanism of RBFox2-driven miR-34a transcriptional repression not fully defined
    • Cytoplasmic cancer role rests on correlative tissue data
  14. 2021 High

    Systematized RBFOX2's combinatorial partnerships and extended its reach to alternative polyadenylation, defining how partner proteins broaden its target repertoire.

    Evidence RIP, partner-specific knockdown and RNA-seq parsing CLIP data for hnRNPC/hnRNPM/SRSF1 modes; 3'-end and nanopore sequencing with binding-site mutagenesis for APA

    PMID:34244793 PMID:34731606

    Open questions at the time
    • Stoichiometry and direct vs. indirect partner contacts not all resolved
    • How RBFOX2 chooses splicing vs. APA outcomes at shared sites unknown
  15. 2021 Medium

    Revealed transcription-factor partnerships and antagonists that retarget or oppose RBFOX2 activity in cancer, including ETS factors, EWS-FLI1, and the SON/hnRNPA2B1 complex.

    Evidence Co-IP, RNA-seq, motif-enrichment and CLIP for ERG/FLI1/EWS-FLI1; SON–hnRNPA2B1 co-IP with RNA-seq and orthotopic GBM xenograft; SLUG-dependent ESRP1 repression assays

    PMID:33089214 PMID:34009296 PMID:34548489

    Open questions at the time
    • Whether interactions are direct or scaffolded not all established
    • Transcriptional (non-RNA-binding) role of RBFOX2 partly inferred
  16. 2023 High

    Defined post-translational control of RBFOX2 abundance and an m6A-reading chromatin-silencing axis, integrating its regulation and its epigenetic output.

    Evidence FBXO7/PRMT5 ubiquitination/methylation site mapping with co-IP and RNA-seq; m6A mapping on caRNAs with RBFOX2–RBM15–YTHDC1 co-IP, H3K27me3 ChIP-seq and AML differentiation assays

    PMID:37640841 PMID:37822160

    Open questions at the time
    • Interplay between stabilizing (FBXO7) and degradative pathways not co-analyzed
    • How m6A recognition is structurally achieved by RBFOX2 unknown
  17. 2023 High

    Established RBFOX2 as a metastasis suppressor acting through cytoskeletal/RHO-GTPase splicing, and as an essential regulator of β-cell insulin secretion.

    Evidence PDX and KO PDA models with in vivo metastasis, RNA-seq and cytoskeletal imaging of MPRIP and ABI1 isoforms; pancreatic conditional KO with glucose physiology, EM granule docking and SNARE-component splicing

    PMID:36949200 PMID:38007492 PMID:38114498

    Open questions at the time
    • Which individual splice events are necessary vs. sufficient for metastasis suppression not fully isolated
    • Direct SNARE-component binding not nucleotide-mapped
  18. 2025 High

    Extended RBFOX2 to stem-cell proteostasis, vascular smooth-muscle ubiquitin regulation/actin control, and continued tissue-specific splicing roles in muscle and heart.

    Evidence Hematopoietic conditional KO with proteostasis assays and chemical rescue; RhoBTB1/CUL3 proximity-labeling MS, co-IP, ubiquitination and SMC-specific KO with arterial stiffness; satellite-cell KO with Numb exon analysis; Snap23 microexon co-regulation with QKI/MBNL1

    PMID:40207498 PMID:41337583 PMID:41418788 PMID:41926228

    Open questions at the time
    • Mechanism linking RBFox2 splicing targets to proteostasis defect unresolved
    • How RhoBTB1/CUL3 degradation integrates with FBXO7 stabilization unclear

Open questions

Synthesis pass · forward-looking unresolved questions
  • How RBFOX2 integrates its multiple activities—cassette splicing, AS-NMD, APA, nuclear PRC2/m6A silencing, and cytoplasmic mRNA regulation—into coordinated, context-specific outputs remains unresolved.
  • No unifying model for how localization, partner availability, and post-translational state select among activities
  • Structural basis for m6A and nascent-RNA recognition undefined
  • Competing stabilizing vs. degradative ubiquitin pathways not reconciled

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0003723 RNA binding 5 GO:0060090 molecular adaptor activity 3 GO:0140110 transcription regulator activity 2
Localization
GO:0005634 nucleus 3 GO:0005829 cytosol 3 GO:0000228 nuclear chromosome 2
Pathway
R-HSA-1266738 Developmental Biology 5 R-HSA-1643685 Disease 4 R-HSA-8953854 Metabolism of RNA 4 R-HSA-74160 Gene expression (Transcription) 2
Complex memberships
PRC2 (nascent-RNA-dependent association)cytoplasmic polyadenylation complex (Xenopus ortholog)cytoplasmic stress granules

Evidence

Reading pass · 50 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2006 RBFOX2 (Fox-2) specifically recognizes the hexameric RNA element UGCAUG, as established by SELEX experiments with Fox-1 (identical RRM to Fox-2). Both Fox-1 and Fox-2 bind UGCAUG elements in the downstream intron of protein 4.1R exon 16 and activate exon inclusion; knockdown of Fox-2 via siRNA decreased exon 16 splicing, identifying Fox-2 as a physiological activator of this splicing switch in differentiating erythroid cells. SELEX, co-transfection splicing assays, siRNA knockdown, immunoblot The Journal of biological chemistry High 16537540
2006 Fox-2 regulates FGFR2 exon IIIb vs. IIIc choice in an epithelial cell-specific manner. Fox-2 expression is higher in IIIb+ epithelial cells; ectopic expression of Fox-2 switches splicing from IIIc to IIIb, absolutely dependent on (U)GCAUG elements. The C-terminal region of Fox-2 is required for this activity. Fox-2 knockdown abolishes the IIIb inclusion switch. Fox-2 also autoregulates its own activity by inducing skipping of its own exon 6 to produce an inactive isoform. RT-PCR splicing assays, Fox-2 overexpression, siRNA knockdown, mutagenesis of UGCAUG elements, isoform analysis Molecular and cellular biology High 16449636
2006 Fox-1 and Fox-2 repress inclusion of calcitonin-specific exon 4 of calcitonin/CGRP pre-mRNA via two UGCAUG elements flanking the 3' splice site. They block U2AF65 binding to the polypyrimidine tract upstream of exon 4, establishing a mechanism of splice-site occlusion. Overexpression in HeLa and neuronal cells, UGCAUG mutation analysis, U2AF65 binding assays Molecular and cellular biology High 17101796
2007 During erythroid differentiation, an erythroid differentiation-inducible Fox-2 isoform (mFox-2A) is upregulated while a commonly expressed isoform (mFox-2F) is downregulated. mFox-2A exerts a much stronger enhancing effect on protein 4.1R exon 16 inclusion than mFox-2F, and isoform-specific knockdown of mFox-2A reverses its splicing-enhancing activity, demonstrating isoform-specific regulation. RT-PCR isoform analysis during erythroid differentiation, isoform-specific overexpression, isoform-specific siRNA knockdown Blood High 17715393
2008 Fox-1/Fox-2 repress calcitonin-specific splicing through a two-step mechanism: (1) binding to an intronic UGCAUG element blocks SF1-dependent E' complex formation; (2) binding to an exonic UGCAUG element blocks the E' to E complex transition. This constitutes a novel strategy of blocking two distinct prespliceosome assembly steps by a single factor. Biochemical spliceosome assembly assays, UGCAUG mutant constructs, gel-shift analyses Molecular and cellular biology High 18573872
2008 Genome-wide prediction and experimental validation established that Fox-1 and Fox-2 bind UGCAUG and that preferred binding position relative to the alternative exon determines splicing outcome: downstream binding activates exon inclusion, whereas upstream binding represses it. Computational binding site analysis, phylogenetic conservation, splicing microarray, experimental validation by RT-PCR Genes & development Medium 18794351
2008 Xenopus Rbm9 (ortholog of RBFOX2/RBM9) is a component of the cytoplasmic polyadenylation complex in Xenopus oocytes; it directly interacts with XGld2 (poly(A) polymerase), is found with CPEB and CPSF, and tethered XRbm9 stimulates translation of a reporter mRNA, revealing a function in cytoplasmic polyadenylation beyond splicing. Co-immunoprecipitation, interaction mapping, tethered-function reporter assay in Xenopus oocytes The FEBS journal Medium 18177378
2010 Fox-2 (RBFOX2) regulates alternative splicing of lysyl hydroxylase 2 (LH2) exon 13A by binding to UGCAUG motifs flanking the exon; overexpression of Fox-2 enhances exon 13A inclusion (generating the scleroderma-associated LH2-long isoform) and knockdown reduces LH2-long mRNA levels in SSc patient fibroblasts. Mutagenesis of two of four Fox-binding motifs is sufficient to abolish exon inclusion. LH2 minigene splicing assays, Fox-2 overexpression/knockdown, UGCAUG mutagenesis, RT-PCR in patient fibroblasts Arthritis and rheumatism High 20131247
2011 RBFOX2 promotes protein 4.1R exon 16 inclusion by recruiting U1 snRNP to the weak 5' splice site via direct interaction between the RBFOX2 C-terminal domain (CTD) and the zinc finger region of U1C protein. Strengthening the 5' splice site to consensus abrogates the requirement for RBFOX2, and CTD deletion abolishes the splicing-enhancing effect. 5' splice site mutagenesis, RBFOX2 CTD deletion analysis, U1C interaction assays, U1 snRNP recruitment assays Molecular and cellular biology High 22083953
2011 Rbfox3 (NeuN) negatively regulates Rbfox2: nuclear Rbfox3 isoforms enhance inclusion of cryptic exons in Rbfox2 mRNA, resulting in nonsense-mediated decay of Rbfox2 transcripts, and Rbfox3 also promotes production of a dominant-negative Rbfox2 protein through alternative splicing of Rbfox2 pre-mRNA. Alternative splicing analysis, RT-PCR, NMD pathway assays, overexpression of Rbfox3 isoforms PloS one Medium 21747913
2012 CNS-specific deletion of Rbfox2 disrupts cerebellar development. Genome-wide analysis identifies Scn8a (Nav1.6 sodium channel) as an Rbfox2 splicing target; improper splicing in Rbfox2-null brains leads to reduced Nav1.6 protein. Combined Purkinje cell deletion of Rbfox1 and Rbfox2 causes irregular firing, establishing that Rbfox2 (with Rbfox1) maintains neuronal pacemaking via sodium channel splicing. Conditional knockout mice, RNA-seq, electrophysiology of Purkinje cells, immunoblot for Nav1.6 Genes & development High 22357600
2012 FOX-2 (RBFOX2) interacts with ataxin-1 (co-immunoprecipitation in mammalian cells) and directly interacts with ataxin-2; reduction of Fox-2 levels leads to increased skipping of ataxin-2 exon 18. Overexpression of nuclear ataxin-1 sequesters Fox-2 into nuclear inclusions and impairs this splicing event. Yeast-2-hybrid, co-immunoprecipitation, siRNA knockdown, RT-PCR splicing assay PloS one Medium 22666429
2013 RBFOX2 expression is upregulated during EMT, correlating with increased RBFOX2-regulated splicing of cortactin, Pard3, and Dnm2. Depletion of RBFOX2 in cells that have completed EMT significantly reduces invasive potential without affecting mesenchymal marker expression or TGF-β signaling, demonstrating a specific role for RBFOX2-regulated splicing in cellular invasion. EMT cell culture models, siRNA knockdown, RT-PCR splicing assays, invasion assays (Matrigel), western blotting Oncogene Medium 23435423
2013 MBNL1 and RBFOX2 cooperatively control a splicing program during mesoderm differentiation. High-throughput RT-PCR during iPSC reprogramming and redifferentiation identified concerted splicing changes in at least 10 conserved genes (including PLOD2, CLSTN1, ATP2A1) controlled by these two splicing regulators. High-throughput RT-PCR during iPSC reprogramming/differentiation, RBP knockdown in cell lines Nature communications Medium 24048253
2014 Rbfox2 cross-regulates AS-coupled NMD (AS-NMD) events within RNA-binding protein genes to alter their steady-state expression. Using iCLIP and RNA-seq in mouse ESCs, >200 AS-NMD events were identified as Rbfox2-bound; these 'silent' events show minimal splicing change but significant gene expression changes upon knockdown due to NMD isoform regulation. Nearly 70 events fall within RBP genes, many of which are autoregulated, establishing Rbfox2 as a regulator of a broader RBP network. iCLIP-seq, RNA-seq, siRNA knockdown, RT-PCR validation of NMD isoforms Genes & development High 24637117
2014 Rbfox2 regulates ~30% of splicing transitions during myogenesis and is specifically required for myoblast fusion. Rbfox2 iCLIP combined with RNA-seq identified Mef2d and Rock2 as direct splicing targets; rescue of myoblast fusion defects in Rbfox2-depleted cultures by restored Mef2d and Rock2 activities demonstrates functional cooperation between isoforms generated by coordinated alternative splicing. iCLIP-seq, RNA-seq, shRNA knockdown, rescue experiments with Mef2d/Rock2 isoforms, fusion assays Molecular cell High 25087874
2015 Transverse aortic constriction (TAC) potently decreases RBFox2 protein in mouse heart. Cardiac-specific ablation of RBFox2 generates phenotypes resembling cardiac decompensation, and RBFox2 regulates splicing of genes involved in heart function. A subset of these genes undergo developmental regulation during postnatal heart remodeling that is reversed in both TAC-treated and RBFox2 knockout mice, establishing RBFox2 as a stress-sensor splicing regulator in pressure overload-induced heart failure. Conditional cardiac knockout mice, TAC surgery model, RNA-seq splicing analysis, RT-PCR validation, echocardiography Cell reports High 25753418
2016 RBFox2 directly interacts with Polycomb Repressive Complex 2 (PRC2) in a nascent RNA-dependent manner. RBFox2 depletion causes widespread increase in nascent RNA production and eradicates PRC2 targeting on the majority of bivalent gene promoters, leading to transcriptional de-repression. ChIP-seq reveals extensive RBFox2 chromatin association dependent on nascent RNA. ChIP-seq, biochemical interaction assays (RBFox2–PRC2 co-IP), nascent RNA sequencing, Bayesian network analysis, RBFox2 depletion in multiple cell types Molecular cell High 27211866
2016 RBFOX2 protein levels are elevated in diabetic hearts due to upregulation of a dominant-negative (DN) isoform of RBFOX2. DN RBFOX2 interacts with wild-type RBFOX2 (co-IP) and inhibits RBFOX2-mediated alternative splicing of target genes. Ectopic expression of DN RBFOX2 impairs intracellular calcium release in cardiomyocytes, identifying dysregulation by DN isoform as an early event in diabetic cardiomyopathy. Co-IP (DN-WT interaction), RT-PCR for splicing, overexpression of DN isoform, calcium imaging in cardiomyocytes, diabetic mouse hearts Cell reports High 27239029
2016 A nonsense mutation in Rbfox2 identified in hypoplastic left heart syndrome (HLHS) patients truncates the protein and impairs its subcellular distribution (nuclear export/localization) and function in RNA metabolism. Rbfox2 regulates mRNA levels of targets through 3'UTR binding sites in addition to splicing regulation. Truncation mutant analysis, subcellular fractionation, RNA-immunoprecipitation, transcriptome analysis of HLHS patient right ventricles Scientific reports Medium 27485310
2016 A nuclear localization signal (NLS) at the N terminus of Rbfox2 isoform 1A (not present in isoform 1F) is identified; Rbfox2 1A isoforms lacking the C-terminal NLS remain nuclear, whereas equivalent 1F isoforms are cytoplasmic. A shift toward cytoplasmic 1F isoforms occurs during EMT. Isoform cloning, subcellular fractionation, immunofluorescence, EMT cell models FEBS letters Medium 27859055
2017 Rbfox2 represses inclusion of alternatively spliced exon 18a in CaV2.2 (Cacna1b) pre-mRNA. siRNA knockdown of Rbfox2 in neuronal cells increases e18a inclusion, and Rbfox2-RNA immunoprecipitation demonstrates reduced Rbfox2 binding upstream of e18a in adult sympathetic neurons. CaV2.2 currents are larger when exon 18a is included, establishing that Rbfox2 limits CaV2.2 current size early in development. siRNA knockdown, RNA immunoprecipitation (RIP)-qPCR, whole-cell patch clamp, RT-PCR eNeuro Medium 29067356
2017 Rbfox2 dynamically regulates alternative exons 9* and 33 of the vascular CaV1.2 (Cacna1c) calcium channel. Knockdown of Rbfox2 in vascular smooth muscle cells increases exon 9* and decreases exon 33 inclusion, shifting CaV1.2 window current to more negative potentials and increasing pressure-induced myogenic tone in mesenteric artery. siRNA knockdown in VSMCs, RT-PCR splicing assays, patch clamp electrophysiology, pressure myograph for myogenic tone Hypertension Medium 28993448
2017 Rbfox2 is a novel constituent of cytoplasmic stress granules. RNA-binding activity of Rbfox is required for its localization into stress granules. In stress granules, Rbfox2 binds and regulates retinoblastoma 1 (RB1) mRNA and protein expression during and after stress exposure. Immunofluorescence co-localization, RNA-immunoprecipitation sequencing (RIP-seq) of cytoplasmic targets, stress granule markers, stress induction assays Scientific reports Medium 28894257
2018 Endothelial-specific deletion of Rbfox2 (the only Rbfox family member expressed in arterial endothelium) suppresses a subset of alternative splicing and transcriptional changes induced by low arterial flow, identifying an Rbfox2-dependent alternative splicing program activated during flow-driven vascular inflammation involving platelet and macrophage recruitment. Endothelial conditional Rbfox2 knockout mice, in vivo low-flow arterial model, RNA-seq, platelet/macrophage depletion eLife High 29293084
2019 Rbfox2 present in cytoplasmic stress granules inhibits RB1 protein expression and promotes cell cycle progression. Resveratrol-induced dissociation of Rbfox2 from stress granules inhibits RB1 suppression and cancer progression; Rbfox2 cytoplasmic localization in colon cancer tissues correlates with its stress granule role distinct from nuclear splicing function. Immunofluorescence in human cancer tissues, stress granule induction/dissolution assays, RB1 protein measurement, tumor growth assays Experimental & molecular medicine Medium 31028247
2019 Neural crest-specific deletion of Rbfox2 causes cleft palate and craniofacial bone defects. Rbfox2 regulates splicing of genes in TGF-β-Tak1 signaling; restoration of TGF-β signaling by Tak1 overexpression rescues proliferation defects in Rbfox2 mutant neural crest cells. A positive feedback loop exists where TGF-β signaling promotes Rbfox2 expression in NCCs. Conditional Rbfox2 knockout in neural crest cells (Wnt1-Cre), RNA-seq, Tak1 overexpression rescue experiments, craniofacial morphology analysis eLife High 31241461
2019 Alternative splicing of exon 10 in RBFOX2 removes a nuclear localization signal, causing cytoplasmic localization of RBFOX2 in calcific tendons vs. nuclear localization in normal tendons. Cytoplasmic RBFOX2 in calcific tendons is associated with altered splicing of RBFOX2 target genes CHD2 and MBNL1. Immunofluorescence microscopy, RBFOX2 isoform sequencing, RT-PCR of splicing targets in clinical samples Experimental and molecular pathology Medium 31128090
2019 RBFox2 controls a feed-forward regulatory pathway in heart failure: reduced RBFox2 induces transcriptional repression of miR-34a, which then targets Jph2 mRNA to impair excitation-contraction coupling, leading to progressive cardiac dysfunction. Administration of miR-34a antagomir alleviates RBFox2 depletion-induced heart dysfunction. Conditional cardiac RBFox2 knockout, miR-34a mimic/antagomir in vivo, Jph2 expression analysis, cardiac function assays (echocardiography) PNAS High 30867288
2020 A non-muscle RBFOX2 isoform (RBFOX240) is upregulated in DM1 heart tissue due to altered splicing factor and microRNA activities. Mice expressing RBFOX240 in heart via transgenic or CRISPR/Cas9 knockin reproduce DM1-related cardiac conduction delay and arrhythmia. RBFOX240 drives splicing defects in voltage-gated sodium and potassium channels, altering their electrophysiological properties. DM1 patient tissue analysis, tetracycline-inducible transgenesis, CRISPR/Cas9 knockin mice, RNA-seq, electrophysiological recording, integration with cardiac transcriptomes Developmental cell High 32109384
2020 RBFOX2 is required for cardiomyocyte alternative splicing regulation during heart development; conditional deletion in embryonic mouse hearts causes cardiac chamber and yolk sac vasculature defects resembling HLHS. RNA-seq identifies dysregulated AS networks affecting cell–ECM adhesion via Rho GTPase cycling genes. Antisense oligos modulating AS of two Rho GTPase cycling genes cause cell cycle and cell-ECM adhesion defects. Cardiac conditional Rbfox2 KO, RNA-seq, antisense oligonucleotide (ASO) experiments, cell adhesion and cycle assays Nucleic acids research High 35137168
2020 RBFOX2 transcriptionally represses ESRP1 by interacting with the transcription factor SLUG under hypoxia/TGF-β signaling, in addition to its RNA-binding role. This leads to skipping of hMENA exon 11a, producing a pro-metastatic isoform. Both RBFOX2 and SLUG are upregulated via TGF-β signaling under hypoxia. TGF-β/hypoxia treatment, ChIP/reporter assays for ESRP1 repression, RBFOX2 and SLUG interaction assays, AS analysis of hMENA NAR cancer Medium 33089214
2021 RBFOX2 can interact with hnRNPC, hnRNPM, and SRSF1 to regulate splicing in three distinct binding modes: (1) single mode—RBFOX2 alone at canonical UGCAUG; (2) multiple mode—RBFOX2 adjacent to at least one other RBP partner; (3) secondary mode—RBFOX2 recruited to sites without its canonical motif through partner binding. These modes control distinct sets of transcripts with different positional relationships to splice sites. RNA immunoprecipitation, RBP-specific knockdown, RNA-seq, splice-sensitive PCR, parsing of public CLIP datasets Nucleic acids research High 34244793
2021 RBFOX2 is critical for maintaining alternative polyadenylation (APA) patterns in myoblasts. 3'-end and nanopore sequencing reveal that RBFOX2 depletion disrupts APA of mitochondrial and contractile genes. Mechanistically, RBFOX2 binding to consensus RBFOX2 motifs near the distal polyadenylation site of Slc25a4 enforces use of the proximal site; RBFOX2 depletion impairs mitochondrial health in myoblasts. 3'-end sequencing, nanopore cDNA sequencing, RBFOX2 knockdown, binding site mutagenesis, mitochondrial function assays Cell reports High 34731606
2021 Rbfox2 mediates exon 11 inclusion in insulin receptor (IR) pre-mRNA in hepatoma cells via an intronic UGCAUG element just downstream of exon 11. Mutation of the UGCAUG sequence causes exon 11 skipping, and Rbfox2 knockdown enhances endogenous exon 11 skipping. SRSF3 binding site mutations combined with Rbfox2 binding site mutations completely abolish exon 11 inclusion. Minigene splicing reporter with deletions and mutations, siRNA knockdown of Rbfox2, RT-PCR Biochimie Medium 34022289
2021 SON suppresses RBFOX2-mediated non-oncogenic neuronal splicing in GBM by forming a complex with hnRNP A2B1 that antagonizes RBFOX2, leading to skipping of RBFOX2-targeted cassette exons including the PTBP2 neuronal exon. SON knockdown restores RBFOX2 activity and inhibits GBM tumor growth in vivo. Co-IP (SON–hnRNP A2B1), RNA-seq after SON knockdown, RT-PCR of RBFOX2 targets, orthotopic xenograft Nature communications Medium 34548489
2021 ERG and FLI1 transcription factors associate with RBFOX2 via their conserved C-terminal domain (ETS domain region), and co-regulate alternative spliced exons (ASEs) enriched in RBFOX2 motifs. EWS-FLI1 also associates with RBFOX2 but antagonizes its effects on exon inclusion, including reducing RBFOX2 binding to ADD3 pre-mRNA, promoting an isoform that represses the mesenchymal phenotype. Co-IP (ERG/FLI1/EWS-FLI1 with RBFOX2), RNA-seq, RBFOX2 motif enrichment analysis, CLIP validation Nucleic acids research Medium 34009296
2022 RBFOX2 promotes inclusion of TEAD1 exon 6 via binding to a conserved GCAUG element in the downstream intron. Full-length TEAD1 (with exon 6) has greater transcriptional activity and YAP interaction than TEAD1ΔE6, with the difference in transcription mediated through YAP binding. This establishes an RBFOX2–TEAD1 splicing axis modulating Hippo–YAP signaling. RT-PCR and RNA-seq for splicing, RBFOX2 knockdown and overexpression, transcriptional activity assays, YAP interaction assays, GCAUG mutagenesis Nucleic acids research Medium 35699208
2022 Liver RBFOX2 regulates cholesterol homeostasis by controlling alternative splicing of Scarb1 (scavenger receptor B1). In diet-induced obesity, RBFOX2 function is decreased in liver causing a Scarb1 isoform switch and altered hepatocyte lipid homeostasis. Splice-switching oligonucleotides targeting this network alleviate obesity-induced liver inflammation and promote anti-atherogenic lipoprotein profile. Enhanced iCLIP in mouse liver, liver-specific Rbfox2 KO, dietary obesity model, ASO treatment, lipoprotein profiling, inflammation assays Nature metabolism High 36536133
2022 RBFOX2 regulates alternative splicing of MICU1 (Mitochondrial Calcium Uniporter activator) during myogenesis, generating a muscle-specific MICU1.1 splice variant with distinct properties for mitochondrial Ca2+ uptake. RBFOX2 is identified as the splicing factor driving this switch during myoblast differentiation. Tissue isoform analysis, myogenesis differentiation time-course, RBFOX2 knockdown, mitochondrial Ca2+ uptake assays International journal of molecular sciences Medium 35269658
2023 RBFOX2 recognizes N6-methyladenosine (m6A) on chromatin-associated RNAs (caRNAs) and recruits RBM15 (an m6A methyltransferase complex component) to facilitate methylation of promoter-associated RNAs. RBM15 interacts with YTHDC1 and recruits PRC2 to RBFOX2-bound loci for chromatin silencing. This RBFOX2/m6A/RBM15/YTHDC1/PRC2 axis is required for AML cell survival and myeloid differentiation control. m6A mapping on caRNAs, Co-IP (RBFOX2–RBM15, RBM15–YTHDC1), ChIP-seq for H3K27me3, RBFOX2 knockdown in AML cells, functional differentiation assays Nature cell biology High 37640841
2023 RBFOX2 acts as a metastatic suppressor in pancreatic ductal adenocarcinoma (PDA). Overexpression of RBFOX2 in metastatic PDA cells reduces metastatic potential in vitro and in vivo; depletion increases it. RBFOX2 splicing targets are enriched in RHO GTPase pathways. Modulation of RBFOX2-regulated splicing of MPRIP (myosin phosphatase RHO-interacting protein) alters cytoskeletal organization and focal adhesion formation. Patient-derived xenograft models, RNA-seq/splicing analysis of RBFOX2 targets, in vivo metastasis assays, cytoskeletal imaging Nature High 36949200
2023 RBFOX2 regulates insulin secretion in pancreatic β-cells through alternative splicing of SNARE complex components required for insulin granule docking and exocytosis. Conditional Rbfox2 mutation in mouse pancreas results in decreased insulin secretion, impaired blood glucose homeostasis, and reduced insulin granule docking. Pancreatic conditional Rbfox2 knockout mice, glucose tolerance/insulin secretion assays, electron microscopy for granule docking, RNA-seq for splicing of SNARE components Nature communications High 38007492
2023 FBXO7 stabilizes Rbfox2 by K63-linked ubiquitination at Lys249 upon arginine dimethylation at Arg341 and Arg441 by PRMT5. This FBXO7-Rbfox2 axis controls splicing of mesenchymal genes (FoxM1, Mta1, Postn) and promotes GBM mesenchymal transformation and chemoresistance. Co-IP, ubiquitination assays, site-specific mutagenesis of Rbfox2 ubiquitination and methylation sites, RNA-seq, tumor xenografts Advanced science Medium 37822160
2023 RBFOX2 depletion promotes pancreatic cancer progression and liver metastasis by altering splicing of cytoskeletal remodeling genes. RBFOX2-mediated splicing of ABI1 (exon 9) controls ABI1 protein isoform abundance and localization; the ABI1 ΔEx9 isoform (generated by splice-switching ASOs) enhances cell migration. Sleeping Beauty insertional mutagenesis screen, RBFOX2 KO in PDAC models, RNA-seq, ABI1 isoform localization by IF, splice-switching ASOs, in vivo metastasis Nature communications High 38114498
2025 Rbfox2 selectively governs hematopoietic stem cell (HSC) self-renewal by regulating proteostasis. Deletion of Rbfox2 from the hematopoietic compartment specifically depletes HSCs but not multipotent progenitors. Mechanistically, Rbfox2 loss leads to increased protein synthesis and accumulated misfolded/unfolded proteins in HSCs. Small molecules restoring proteostasis rescue HSC defects in Rbfox2-deficient mice. Conditional Rbfox2 KO in hematopoietic compartment, bone marrow transplantation, proteostasis assays (protein synthesis rate, unfolded protein content), chemical rescue experiments Science advances High 41337583
2025 RBFOX2 is ubiquitinated in a RhoBTB1- and CUL3-dependent manner in vascular smooth muscle cells, establishing it as a substrate of the RhoBTB1/CUL3 ubiquitin-proteasome pathway. Co-immunoprecipitation validates the RBFOX2–RhoBTB1 interaction. Rbfox2 depletion impairs the actin cytoskeleton and alters filamentous/globular actin levels. SMC-specific Rbfox2 deletion halts progression of ANG-induced arterial stiffness. Proximity labeling + mass spectrometry, co-immunoprecipitation, ubiquitination assays, siRNA knockdown, conditional SMC-specific Rbfox2 KO mice, actin polymerization assays, arterial stiffness measurements JCI insight High 41926228
2025 RBFOX2 is essential for cardiomyocyte differentiation by promoting exon usage shifts to mature patterns in sarcomere and cytoskeletal genes (including ACTN2). RBFOX2 autoregulates itself at mutually exclusive exons: at critical RBFOX2 levels, autoregulation enforces mature isoforms; in heterozygous CMs, autoregulation is disrupted generating a dominant-negative product. Overexpression of ACTN2 rescues heterozygous (not null) phenotypes by restoring contractility and triggering mechanosensing upregulation of RBFOX2 from the wildtype allele. iPSC-derived cardiomyocyte differentiation, RBFOX2 heterozygous and null mutants, RNA-seq, ACTN2 overexpression rescue, contractility assays bioRxivpreprint Medium 41280054
2025 RBFOX2 promotes inclusion of the Numb exon 6 in satellite cells (SCs), and RBFOX2 loss delays SC activation and muscle regeneration. Exon 6-containing Numb is required for SC activation; its skipping upregulates Notch signaling and delays activation. In vivo SC fixation to preserve quiescent state, Rbfox2 conditional KO in SCs, RNA-seq, exon 6 inclusion/skipping analysis, muscle regeneration assay Stem cell reports Medium 41418788
2025 RBFOX2 promotes splicing of a Snap23 microexon in striated muscle by binding downstream of the microexon together with QKI, and this regulation can be escaped when the weak splice donor is mutated to consensus. MBNL1 acts as an additional, minor layer of Snap23 microexon control. The microexon is mis-regulated in mouse models of heart and skeletal muscle diseases. Splicing factor knockdowns (RBFOX2, QKI, MBNL1), minigene splice site mutagenesis, disease model analysis, binding site mapping RNA biology Medium 40207498

Source papers

Stage 0 corpus · 90 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2008 Defining the regulatory network of the tissue-specific splicing factors Fox-1 and Fox-2. Genes & development 259 18794351
2012 The splicing regulator Rbfox2 is required for both cerebellar development and mature motor function. Genes & development 182 22357600
2013 The RNA-binding protein Rbfox2: an essential regulator of EMT-driven alternative splicing and a mediator of cellular invasion. Oncogene 138 23435423
2012 RBFOX2 is an important regulator of mesenchymal tissue-specific splicing in both normal and cancer tissues. Molecular and cellular biology 130 23149937
2014 Rbfox2 controls autoregulation in RNA-binding protein networks. Genes & development 115 24637117
2013 MBNL1 and RBFOX2 cooperate to establish a splicing programme involved in pluripotent stem cell differentiation. Nature communications 112 24048253
2018 The long non-coding RNA MALAT1 promotes ovarian cancer progression by regulating RBFOX2-mediated alternative splicing. Molecular carcinogenesis 103 30294913
2014 Rbfox2-coordinated alternative splicing of Mef2d and Rock2 controls myoblast fusion during myogenesis. Molecular cell 103 25087874
2006 Fox-2 splicing factor binds to a conserved intron motif to promote inclusion of protein 4.1R alternative exon 16. The Journal of biological chemistry 100 16537540
2006 Fox-2 mediates epithelial cell-specific fibroblast growth factor receptor 2 exon choice. Molecular and cellular biology 98 16449636
2011 NeuN/Rbfox3 nuclear and cytoplasmic isoforms differentially regulate alternative splicing and nonsense-mediated decay of Rbfox2. PloS one 88 21747913
2015 Repression of the Central Splicing Regulator RBFox2 Is Functionally Linked to Pressure Overload-Induced Heart Failure. Cell reports 85 25753418
2023 RBFOX2 modulates a metastatic signature of alternative splicing in pancreatic cancer. Nature 75 36949200
2016 RBFox2 Binds Nascent RNA to Globally Regulate Polycomb Complex 2 Targeting in Mammalian Genomes. Molecular cell 71 27211866
2023 RBFOX2 recognizes N6-methyladenosine to suppress transcription and block myeloid leukaemia differentiation. Nature cell biology 70 37640841
2016 Dysregulation of RBFOX2 Is an Early Event in Cardiac Pathogenesis of Diabetes. Cell reports 68 27239029
2022 CircRAPGEF5 interacts with RBFOX2 to confer ferroptosis resistance by modulating alternative splicing of TFRC in endometrial cancer. Redox biology 61 36182807
2013 Aeromonas spp. simultaneously harbouring bla(CTX-M-15), bla(SHV-12), bla(PER-1) and bla(FOX-2), in wild-growing Mediterranean mussel (Mytilus galloprovincialis) from Adriatic Sea, Croatia. International journal of food microbiology 55 23973842
2006 Role for Fox-1/Fox-2 in mediating the neuronal pathway of calcitonin/calcitonin gene-related peptide alternative RNA processing. Molecular and cellular biology 54 17101796
1997 A novel class C beta-lactamase (FOX-2) in Escherichia coli conferring resistance to cephamycins. Antimicrobial agents and chemotherapy 53 9303413
2016 Rbfox2 function in RNA metabolism is impaired in hypoplastic left heart syndrome patient hearts. Scientific reports 52 27485310
2019 Rbfox2 dissociation from stress granules suppresses cancer progression. Experimental & molecular medicine 51 31028247
2020 Hypoxia-induced TGF-β-RBFOX2-ESRP1 axis regulates human MENA alternative splicing and promotes EMT in breast cancer. NAR cancer 49 33089214
2022 RBFOX2 is required for establishing RNA regulatory networks essential for heart development. Nucleic acids research 43 35137168
2019 RBFox2-miR-34a-Jph2 axis contributes to cardiac decompensation during heart failure. Proceedings of the National Academy of Sciences of the United States of America 41 30867288
2017 Stress Granules Contain Rbfox2 with Cell Cycle-related mRNAs. Scientific reports 41 28894257
2020 Aberrant Expression of a Non-muscle RBFOX2 Isoform Triggers Cardiac Conduction Defects in Myotonic Dystrophy. Developmental cell 39 32109384
2021 RBFOX2 alters splicing outcome in distinct binding modes with multiple protein partners. Nucleic acids research 38 34244793
2021 SON drives oncogenic RNA splicing in glioblastoma by regulating PTBP1/PTBP2 switching and RBFOX2 activity. Nature communications 38 34548489
2018 Alternative RNA splicing in the endothelium mediated in part by Rbfox2 regulates the arterial response to low flow. eLife 38 29293084
2007 Regulated Fox-2 isoform expression mediates protein 4.1R splicing during erythroid differentiation. Blood 38 17715393
2011 RBFOX2 promotes protein 4.1R exon 16 selection via U1 snRNP recruitment. Molecular and cellular biology 35 22083953
2017 Aberrant Splicing Induced by Dysregulated Rbfox2 Produces Enhanced Function of CaV1.2 Calcium Channel and Vascular Myogenic Tone in Hypertension. Hypertension (Dallas, Tex. : 1979) 34 28993448
2022 Liver RBFOX2 regulates cholesterol homeostasis via Scarb1 alternative splicing in mice. Nature metabolism 33 36536133
2019 Neural crest-specific deletion of Rbfox2 in mice leads to craniofacial abnormalities including cleft palate. eLife 33 31241461
2023 RBFOX2 deregulation promotes pancreatic cancer progression and metastasis through alternative splicing. Nature communications 31 38114498
2021 RBFOX2 is critical for maintaining alternative polyadenylation patterns and mitochondrial health in rat myoblasts. Cell reports 29 34731606
2021 The oncogenic kinase NEK2 regulates an RBFOX2-dependent pro-mesenchymal splicing program in triple-negative breast cancer cells. Journal of experimental & clinical cancer research : CR 29 34930366
2021 Alternative microexon splicing by RBFOX2 and PTBP1 is associated with metastasis in colorectal cancer. International journal of cancer 27 34346508
1995 Molecular cloning, sequencing and sequence analysis of the fox-2 gene of Neurospora crassa encoding the multifunctional beta-oxidation protein. Molecular & general genetics : MGG 26 7715608
2022 RBFOX2-regulated TEAD1 alternative splicing plays a pivotal role in Hippo-YAP signaling. Nucleic acids research 25 35699208
2021 RBFOX2/GOLIM4 Splicing Axis Activates Vesicular Transport Pathway to Promote Nasopharyngeal Carcinogenesis. Advanced science (Weinheim, Baden-Wurttemberg, Germany) 25 34180133
2008 Repression of prespliceosome complex formation at two distinct steps by Fox-1/Fox-2 proteins. Molecular and cellular biology 25 18573872
2023 Hypoxia induces hepatocellular carcinoma metastasis via the HIF-1α/METTL16/lnc-CSMD1-7/RBFOX2 axis. iScience 24 38089592
2021 ERG transcription factors have a splicing regulatory function involving RBFOX2 that is altered in the EWS-FLI1 oncogenic fusion. Nucleic acids research 23 34009296
2014 RBFOX2 protein domains and cellular activities. Biochemical Society transactions 22 25110022
2016 RBFOX1 and RBFOX2 are dispensable in iPSCs and iPSC-derived neurons and do not contribute to neural-specific paternal UBE3A silencing. Scientific reports 19 27146458
2023 FBXO7 Confers Mesenchymal Properties and Chemoresistance in Glioblastoma by Controlling Rbfox2-Mediated Alternative Splicing. Advanced science (Weinheim, Baden-Wurttemberg, Germany) 18 37822160
2022 LncRNA MIAT Promotes Spinal Cord Injury Recovery in Rats by Regulating RBFOX2-Mediated Alternative Splicing of MCL-1. Molecular neurobiology 17 35641779
2023 Dysregulated Rbfox2 produces aberrant splicing of CaV1.2 calcium channel in diabetes-induced cardiac hypertrophy. Cardiovascular diabetology 16 37415128
2023 Modulation of insulin secretion by RBFOX2-mediated alternative splicing. Nature communications 16 38007492
2019 Genes of the cGMP-PKG-Ca2+ signaling pathway are alternatively spliced in cardiomyopathy: Role of RBFOX2. Biochimica et biophysica acta. Molecular basis of disease 15 31778749
2022 p53-Induced LINC00893 Regulates RBFOX2 Stability to Suppress Gastric Cancer Progression. Frontiers in cell and developmental biology 14 35127712
2017 Cell-Specific RNA Binding Protein Rbfox2 Regulates CaV2.2 mRNA Exon Composition and CaV2.2 Current Size. eNeuro 14 29067356
2010 Fox-2 protein regulates the alternative splicing of scleroderma-associated lysyl hydroxylase 2 messenger RNA. Arthritis and rheumatism 14 20131247
2022 LncRNA ZFAS1 promotes laryngeal cancer progression through RBFOX2-mediated MENA alternative splicing. Environmental toxicology 13 36336961
2019 Alternative splicing induces cytoplasmic localization of RBFOX2 protein in calcific tendinopathy. Experimental and molecular pathology 13 31128090
2012 Associations of polymorphisms in the genes of FGFR2, FGF1, and RBFOX2 with breast cancer risk by estrogen/progesterone receptor status. Molecular carcinogenesis 13 23143756
2011 Evidence that "brain-specific" FOX-1, FOX-2, and nPTB alternatively spliced isoforms are produced in the lens. Current eye research 12 21714144
2008 Xenopus Rbm9 is a novel interactor of XGld2 in the cytoplasmic polyadenylation complex. The FEBS journal 11 18177378
2022 The Splicing of the Mitochondrial Calcium Uniporter Genuine Activator MICU1 Is Driven by RBFOX2 Splicing Factor during Myogenic Differentiation. International journal of molecular sciences 10 35269658
2020 The effect of Rbfox2 modulation on retinal transcriptome and visual function. Scientific reports 10 33184471
2024 Gastrin-related circRNA_0017065 promotes the proliferation and metastasis of colorectal cancer through the miR-3174/RBFOX2 axis. Biology direct 8 39198845
2023 Suppression of RBFox2 by Multiple MiRNAs in Pressure Overload-Induced Heart Failure. International journal of molecular sciences 8 36674797
2022 Alternative Splicing of the Aryl Hydrocarbon Receptor Nuclear Translocator (ARNT) Is Regulated by RBFOX2 in Lymphoid Malignancies. Molecular and cellular biology 8 35404107
2019 A Small Cyclic β-Hairpin Peptide Mimics the Rbfox2 RNA Recognition Motif and Binds to the Precursor miRNA 20b. Chembiochem : a European journal of chemical biology 8 30537200
2016 Identification of a classic nuclear localization signal at the N terminus that regulates the subcellular localization of Rbfox2 isoforms during differentiation of NMuMG and P19 cells. FEBS letters 8 27859055
2023 RBFOX2 regulated EYA3 isoforms partner with SIX4 or ZBTB1 to control transcription during myogenesis. iScience 7 38026174
2020 Trophoblast lineage specific expression of the alternative splicing factor RBFOX2 suggests a role in placental development. Placenta 7 32762877
2012 FOX-2 dependent splicing of ataxin-2 transcript is affected by ataxin-1 overexpression. PloS one 7 22666429
2021 Rbfox2 mediates exon 11 inclusion in insulin receptor pre-mRNA splicing in hepatoma cells. Biochimie 6 34022289
2024 RNA splicing factor RBFOX2 is a key factor in the progression of cancer and cardiomyopathy. Clinical and translational medicine 5 39243148
2017 Novel Rbfox2 isoforms associated with alternative exon usage in rat cortex and suprachiasmatic nucleus. Scientific reports 5 28855650
2024 A novel m6A reader RBFOX2 expression is increased in oral squamous cell carcinoma and promotes tumorigenesis. Journal of stomatology, oral and maxillofacial surgery 4 39244024
2023 RBFOX2 confers tumor growth by PI3K/AKT and MAPK signaling in gastric cancer. European journal of cancer prevention : the official journal of the European Cancer Prevention Organisation (ECP) 4 37264873
2025 Upregulation of WDR4 mediated by RBFOX2 promotes laryngeal cancer progression through the WDR4/m7G/lncRNA ZFAS1/RBFOX2 axis. Naunyn-Schmiedeberg's archives of pharmacology 3 39774908
2024 RBFOX2 as a regulatory linchpin in cancer: insights from a comprehensive review of its roles in tumorigenesis. American journal of cancer research 3 39553227
2022 Non-canonical splice junction processing increases the diversity of RBFOX2 splicing isoforms. The international journal of biochemistry & cell biology 3 35124219
2025 Alternative splicing of the Snap23 microexon is regulated by MBNL, QKI, and RBFOX2 in a tissue-specific manner and is altered in striated muscle diseases. RNA biology 2 40207498
2025 RBFOX2 induces osteogenic differentiation by Jph2 expression in MC3T3-E1 preosteoblast cells. Molecular biology reports 1 40232550
2025 RBFOX2-dependent alternative splicing of Numb regulates Notch signaling during muscle stem cell activation. Stem cell reports 1 41418788
2021 Expression of the alternative splicing regulator Rbfox2 during placental development is differentially regulated in preeclampsia mouse models. American journal of reproductive immunology (New York, N.Y. : 1989) 1 34363260
2026 Vascular smooth muscle RbFox2 regulates the cytoskeleton and arterial stiffness by a RhoBTB1/Cullin-3 mechanism. JCI insight 0 41926228
2026 The RNA-binding protein RBFOX2 suppresses colorectal cancer proliferation and metastasis by reducing FUBP1 mRNA stability to induce mitochondrial dysfunction and ferroptosis. Discover oncology 0 42223720
2025 Genome Sequencing Identifies a Heterozygous Deletion of RBFOX2 in a Family With Congenital Heart Disease: A Case Report. American journal of medical genetics. Part A 0 41159281
2025 RBFOX2: An RNA-binding protein with alternative splicing and non-alternative splicing regulatory functions. Experimental and molecular pathology 0 41172617
2025 Dosage-sensitive RBFOX2 autoregulation promotes cardiomyocyte differentiation by maturing the transcriptome. bioRxiv : the preprint server for biology 0 41280054
2025 Rbfox2 selectively governs hematopoietic stem cell self-renewal by regulating proteostasis. Science advances 0 41337583
2025 An RBPMS-driven splicing regulatory axis including MBNL1, RBFOX2, and QK promotes smooth muscle cell contractile identity. Nucleic acids research 0 41428739
2023 [Loss of RBFOX2 inhibits meiotic initiation in male mice]. Sheng wu gong cheng xue bao = Chinese journal of biotechnology 0 37877394

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