Affinage

SRA1

Steroid receptor RNA activator 1 · UniProt Q9HD15

Round 2 corrected
Length
224 aa
Mass
24.4 kDa
Annotated
2026-04-28
130 papers in source corpus 28 papers cited in narrative 28 extracted findings

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

Insufficient on-target evidence to synthesize a narrative — discovery timeline does not match the canonical SRA1 protein.

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
No controlled-vocabulary terms were assigned to this entry.

Evidence

Reading pass · 28 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
1999 SRA (Steroid Receptor RNA Activator) was identified as a novel transcriptional coactivator that functions as an RNA transcript rather than a protein; it is selective for steroid hormone receptors, mediates transactivation via their amino-terminal AF-1 activation function, and exists in distinct ribonucleoprotein complexes including one containing SRC-1. Evidence that SRA functions as RNA: coactivation persists in the presence of cycloheximide and SRA mutants with multiple translational stop signals retain coactivator activity. Biochemical fractionation, ribonucleoprotein complex isolation, functional reporter assays with cycloheximide treatment, translational stop-signal mutagenesis Cell High 10199399
2001 The RNA-binding DEAD-box proteins p72/p68 function as estrogen receptor alpha (ERα) coactivators through the AF-1 domain by associating with a coactivator complex containing SRA and SRC-1/TIF2. p72/p68 interact with the AD2 domain of SRC-1/TIF2 family proteins and the ERα A/B domain, and co-immunoprecipitate with estrogen-bound ERα in MCF7 cells. Combination of p72/p68 with SRA and TIF2 produces synergistic estrogen-induced transactivation of ERα. Co-immunoprecipitation from MCF7 cells and HeLa nuclear extracts, transient transfection reporter assays, immunofluorescence co-localization The EMBO journal High 11250900
2003 SRA coactivates ERα in a ligand-independent manner through the AF-1 domain, requiring an intact serine-118 (S118) residue in ERα AF-1. MAPK activation induces ligand-independent coactivation of ERα by SRA; activated H-Ras (H-RasV12) cannot rescue S118 ERα mutant activity, indicating that MAPK-mediated phosphorylation of S118 participates in SRA's AF-1-dependent effect. SRA does not coactivate ERβ via AF-1 in the same manner, demonstrating isoform selectivity. Transient transfection reporter assays, site-directed mutagenesis of ERα S118, co-expression with activated Ras and MAPK pathway components The Journal of steroid biochemistry and molecular biology Medium 12943696
2003 In Drosophila cells, Sra-1 (ortholog of human CYFIP1/SRA1 WAVE complex subunit), together with Abi and Kette (Nap1), protects SCAR/WAVE from proteasome-mediated degradation and is critical for SCAR localization and for Arp2/3-dependent actin protrusion generation. RNAi-mediated removal of Sra-1, Abi, or Kette abolishes dynamic protrusions, demonstrating these proteins are essential components of the SCAR regulatory complex downstream of Rac and Cdc42. RNAi screen in Drosophila adherent cells, immunofluorescence, proteasome inhibitor experiments, epistasis analysis Current biology : CB High 14588242
2004 Sra-1 (CYFIP/PIR121) and Nap1 are constitutive components of a WAVE2/Abi-1-containing complex that links Rac1 signaling to site-directed actin assembly and lamellipodia formation. RNA interference removal of Sra-1 or Nap1 abrogates Rac-dependent lamellipodia induced by growth factor stimulation or aluminum fluoride treatment, and microinjection of constitutively active Rac fails to restore lamellipodia in cells lacking either protein. Sra-1, Nap1, WAVE2, and Abi-1 translocate to tips of membrane protrusions after active Rac microinjection. RNAi knockdown, microinjection of constitutively active Rac1, immunofluorescence, co-immunoprecipitation of complex components The EMBO journal High 14765121
2004 The Arabidopsis PIROGI gene encodes a functional ortholog of the human WAVE complex subunit SRA1 (CYFIP1). PIROGI/SRA1 directly interacts with the small GTPase ATROP2 with isoform specificity and selectivity for active (GTP-bound) forms. Both PIROGI and human SRA1 are functionally interchangeable and display identical physical interactions with RHO family GTPases and the WAVE complex subunit NAP125, indicating deeply conserved molecular function in ARP2/3-dependent actin regulation. Genetic complementation, yeast two-hybrid, pull-down assays with active/inactive GTPase forms, phenotypic analysis of pir mutants Development (Cambridge, England) High 15294869
2004 Drosophila Sra-1 interacts physically with Kette (Nap1) in vivo, and both proteins stabilize each other. Sra-1 is expressed in neuronal growth cones; RNAi knockdown of sra-1 in photoreceptor neurons causes stalling of axonal growth, and knockdown in motoneurons produces abnormal neuromuscular junctions similar to kette hypomorphic mutants. A membrane-bound Sra-1 lacking the Kette-binding domain phenocopies loss of function, indicating Sra-1 function is mediated through Kette and controls F-actin cytoskeleton in a Wasp-dependent manner. In vivo co-immunoprecipitation, RNAi in specific neuronal subtypes, dominant-negative Sra-1 construct lacking Kette-binding domain, confocal imaging of neuromuscular junctions Development (Cambridge, England) High 15269173
2005 CRMP-2 interacts with the Sra-1/WAVE1 complex and with the light chain of kinesin-1, linking kinesin-1 to the Sra-1/WAVE1 complex for anterograde transport to axonal growth cones. Knockdown of Sra-1 and WAVE1 cancels CRMP-2-induced axon outgrowth and multiple-axon formation in hippocampal neurons; knockdown of CRMP-2 or kinesin-1 delocalizes Sra-1 and WAVE1 from growth cones, establishing a kinesin-1-dependent transport mechanism for the Sra-1/WAVE1 complex in axon formation. Co-immunoprecipitation, RNAi knockdown in hippocampal neurons, immunofluorescence localization in growth cones Molecular and cellular biology High 16260607
2006 The RNA helicases p68/p72 (DDX5/DDX17) are MyoD-associated proteins, and the non-coding RNA SRA co-immunoprecipitates with MyoD. Both p68/p72 and SRA function as coactivators of MyoD in skeletal muscle differentiation; RNAi knockdown of either p68/p72 or SRA prevents activation of muscle gene expression and cell differentiation. Loss of p68/p72 impairs recruitment of TBP, RNA polymerase II, and the SWI/SNF catalytic subunit Brg-1, and hinders chromatin remodeling at muscle gene promoters. Co-immunoprecipitation with MyoD, RNAi knockdown, chromatin immunoprecipitation, reporter assays, differentiation assays Developmental cell High 17011493
2006 SLIRP (SRA stem-loop interacting RNA binding protein) binds a functional substructure of SRA RNA (STR7) via its RNA recognition motif (RRM) and functions as a nuclear receptor corepressor. SLIRP represses nuclear receptor transactivation in a SRA- and RRM-dependent manner, augments tamoxifen's effect, modulates SRC-1 association with SRA, and is recruited to endogenous promoters in a SRA-dependent manner. SHARP also binds STR7, augmenting repression with SLIRP. RNA pull-down, co-immunoprecipitation, reporter assays, chromatin immunoprecipitation, RRM mutagenesis, immunofluorescence Molecular cell High 16762838
2006 Alternative splicing of the first intron of SRA1 generates both coding (SRAP-producing) and non-coding SRA RNA isoforms in breast cancer cell lines. The intron-1 sequence is sufficient to drive differential splicing in both endogenous and minigene contexts. The relative proportion of coding vs. non-coding SRA transcripts varies across breast cancer cell lines, suggesting that alternative splicing is a regulatory mechanism controlling the balance between SRA RNA coactivator and SRAP protein. 5'-RACE, RT-PCR with isoform-specific primers, minigene splicing assay in breast cancer cell lines DNA and cell biology Medium 16848684
2007 SRA1 encodes both a non-coding RNA (SRA) and an endogenous protein (SRAP) through alternative splicing of intron-1; retention of intron-1 disrupts the SRAP open reading frame, generating the non-coding isoform. Both SRA RNA and SRAP have opposing regulatory roles on steroid receptor signaling and are conserved among Chordata, establishing SRA1 as a bifaceted gene. Review/synthesis of published molecular characterization; original data on isoform identification referenced Nuclear receptor signaling Low 17710122
2008 SRA1 gene knockout (SRA-/-) in zebrafish using morpholino antisense reagents results in a phenotype of myocardial contractile dysfunction, demonstrating that SRA1 (along with HBEGF and IK, which cosegregate in a 600-kb linkage disequilibrium block) independently contributes to cardiac muscle function in vivo. Morpholino antisense knockdown in zebrafish, cardiac function assessment Genome research Medium 19064678
2009 SRA functions as an RNA coactivator for steroidogenic factor 1 (SF-1), and both SF-1 and Dax-1 bind SRA. Dax-1 acts as a dosage-dependent SF-1 coactivator through SRA; SRA knockdown abolishes Dax-1-mediated coactivation and reduces expression of steroidogenic gene products (StAR, MC2R) in adrenal Y1 cells. TIF2 associates with Dax-1 and synergistically coactivates SF-1 target gene transcription in a SRA-dependent manner. Co-immunoprecipitation, siRNA knockdown of SRA, reporter assays, qRT-PCR of steroidogenic gene products Molecular and cellular biology High 19188450
2010 SRA non-coding RNA enhances myogenic differentiation and myogenic conversion through co-activation of MyoD activity, while SRAP (the protein encoded by the same locus) counteracts this SRA RNA-dependent co-activation. SRAP's inhibitory effect is mediated through its RRM-like domain interacting with functional sub-structure STR7 of SRA RNA, establishing a regulatory feedback between the two products of the SRA1 locus. Constructs producing exclusively SRA ncRNA or SRAP protein, myogenic differentiation assays in human satellite cells, co-immunoprecipitation, domain mutagenesis Nucleic acids research High 20855289
2010 SRA non-coding RNA associates with PPARγ and coactivates PPARγ-dependent gene expression to promote adipogenesis. SRA overexpression in ST2 mesenchymal precursor cells promotes adipocyte differentiation; SRA knockdown inhibits 3T3-L1 preadipocyte differentiation. SRA also increases glucose uptake and Akt/FOXO1 phosphorylation in response to insulin, promotes S-phase entry by decreasing p21Cip1 and p27Kip1 expression, and inhibits TNFα-induced JNK phosphorylation, demonstrating multiple signaling roles. Co-immunoprecipitation of SRA with PPARγ, reporter assays, siRNA knockdown, overexpression in ST2 cells, flow cytometry, Western blotting for signaling pathway components PloS one High 21152033
2010 The DEAD-box RNA helicase p68 (DDX5) and its associated non-coding RNA SRA form a complex with CTCF that is essential for insulator function at the IGF2/H19 imprinted control region (ICR). Depletion of SRA or p68 reduces CTCF-mediated insulator activity, increases IGF2 expression, and reduces cohesin binding to CTCF sites without affecting CTCF binding itself. p68/SRA interacts with cohesin complex members, suggesting that p68/SRA stabilizes cohesin-CTCF interaction. ChIP, co-immunoprecipitation, siRNA knockdown, chromosome conformation capture (3C), reporter assays for insulator function Genes & development High 20966046
2010 Dax1 augments LRH-1-mediated activation of the Oct4 gene in mouse embryonic stem cells through a mechanism dependent on the RNA coactivator SRA; SRA knockdown abolishes Dax1-mediated coactivation of Oct4. Dax1 and LRH-1 co-localize at 43% of LRH-1 target genes involved in mES cell pluripotency, and SRA is required for their co-activation. Co-immunoprecipitation of Dax1 with LRH-1, ChIP at Oct4 promoter, luciferase reporter assays, siRNA knockdown of SRA, genome-wide ChIP analysis Molecular endocrinology (Baltimore, Md.) Medium 20943815
2010 Protein complexes containing CYFIP/Sra/PIR121 coordinate Arf1 and Rac1 signaling during clathrin-AP-1-coated carrier biogenesis at the trans-Golgi network (TGN). CYFIP-containing complexes are recruited to TGN membranes by Arf1 GTPase; Rac1 and its exchange factor β-PIX then activate these complexes to promote N-WASP- and Arp2/3-dependent actin polymerization and tubule formation. These events were reconstituted with synthetic membranes. Immunoprecipitation of CYFIP complexes, synthetic membrane reconstitution assay, siRNA knockdown, fluorescence microscopy, dominant-negative GTPase experiments Nature cell biology High 20228810
2010 SRA RNA depletion in human cancer cell lines (MCF-7 and MDA-MB-231) by siRNA reveals hundreds of endogenous SRA target genes, with the majority reduced upon knockdown, implicating SRA as a broad endogenous coactivator. Novel SRA targets identified include SLC2A3, SLC2A12, CCL20, TGFB2, DIO2, TMEM65, TBL1X, and TMPRSS2. SRA depletion in MDA-MB-231 cells reduces invasiveness. siRNA knockdown, microarray gene expression profiling, reporter assays for specific promoters Molecular endocrinology (Baltimore, Md.) Medium 20219889
2013 The cytoplasmic RISC proteins PACT, TRBP, and Dicer bind SRA RNA and function as nuclear receptor coregulators. These RISC proteins, together with Argonaute 2, associate with SRA and specific pre-microRNAs in both the nucleus and cytoplasm and target steroid-responsive promoters to regulate nuclear receptor activity and downstream gene expression. RNA pull-down with SRA, co-immunoprecipitation, ChIP at steroid-responsive promoters, reporter assays, immunofluorescence showing nuclear/cytoplasmic localization Proceedings of the National Academy of Sciences of the United States of America Medium 23550157
2013 RNA helicase Ddx5 (p68) is a novel component of the RBP-J/NICD (Notch intracellular domain) complex; Ddx5 directly interacts with RBP-J and localizes to RBP-J binding sites at Notch target gene loci (preTCRα, Hes1, CD25) in a Notch-dependent manner. SRA, as a cofactor of Ddx5, is also required for Notch target gene activation; knockdown of SRA downregulates Hes1 and preTCRα, associated with loss of p300 occupancy and decreased histone acetylation at Notch target genes. Biotinylation-tagging/mass spectrometry, Co-IP, ChIP, siRNA knockdown, reporter assays Biochimica et biophysica acta Medium 23396200
2014 Whole-body Sra1 gene knockout (SRA-/-) mice are resistant to high-fat diet-induced obesity with decreased fat mass, increased insulin sensitivity (reduced fasting insulin, improved glucose tolerance), decreased adipocyte marker gene expression, reduced plasma TNFα, and fewer hepatic lipid droplets with decreased lipogenesis-associated gene expression. This demonstrates an in vivo role for the SRA1 gene product in adipose tissue mass and glucose homeostasis. Gene knockout mouse model (whole-body Sra1 KO), high-fat diet feeding, body composition analysis, glucose/insulin tolerance tests, gene expression analysis, histology The Journal of biological chemistry High 24675075
2014 The crystal structure of the carboxy-terminal domain of human SRAP (SRA protein) reveals a five-helix bundle that is structurally distinct from known RNA-binding motifs and instead resembles the C-terminal domain of the yeast spliceosome protein PRP18. The structure does not contain the postulated RRM (RNA recognition motif), and direct RNA-binding experiments (in vitro with recombinant SRAP and in cell-based assays) failed to reveal a specific interaction between SRAP and SRA RNA, necessitating re-evaluation of the current model for SRAP function. Crystal structure determination, RNA-binding assays in vitro and in cells, siRNA knockdown with RNA-seq, immunofluorescence for subcellular localization Journal of molecular biology High 24486609
2015 SRA lncRNA participates in transcriptional regulation through complex formation with both trithorax group (TrxG) and polycomb repressive complex 2 (PRC2) complexes. The SRA-associated RNA helicase p68 preferentially stabilizes SRA-TrxG complex formation over SRA-PRC2. SRA binding sites co-occupied by p68 are enriched for H3K4me3 in NTERA2 pluripotent stem cells. NANOG directly interacts with SRA and co-localizes with it genome-wide. SRA is required for maintaining the stem cell state and for reprogramming human fibroblasts to pluripotency. Co-immunoprecipitation of SRA with TrxG and PRC2 components, ChIP-seq for histone marks, RNA immunoprecipitation, genome-wide SRA binding analysis, reprogramming assays PLoS genetics Medium 26496121
2016 LncRNA SRA promotes hepatic steatosis by repressing expression of adipose triglyceride lipase (ATGL). SRA inhibits ATGL promoter activity primarily by inhibiting the inductive effects of the transcription factor FoxO1. Loss of SRA in primary hepatocytes or hepatocyte cell lines upregulates ATGL expression and free fatty acid β-oxidation, while forced SRA expression inhibits ATGL and β-oxidation. Hepatic SRA and ATGL levels are inversely regulated by fasting in mice. Sra KO mice, primary hepatocyte isolation, siRNA and overexpression in hepatocyte cell line, ATGL promoter-luciferase reporter assays, FoxO1 co-transfection experiments Scientific reports Medium 27759039
2016 In ovarian endometriotic stromal cells (ESCs), SRA1 silencing via siRNA increases ERα expression but reduces ERβ expression, and attenuates proliferation while promoting early apoptosis. LncRNA SRA and ERα are expressed at lower levels, while SRAP and ERβ are higher, in ovarian endometriotic tissues versus normal endometrial tissues, establishing SRA1 products as regulators of estrogen receptor isoform balance in endometriosis. siRNA knockdown in primary ESCs, RT-PCR and Western blotting for ER isoforms, CCK-8 proliferation assay, flow cytometry for apoptosis, IHC on tissue specimens Reproductive sciences (Thousand Oaks, Calif.) Low 27694140
2021 SRSF1 promotes inclusion of exon 3 of SRA1 pre-mRNA to generate the long isoform SRA1-L (which promotes HCC cell invasion) by directly binding to exon 3 of SRA1 pre-mRNA. RNA immunoprecipitation, cross-link immunoprecipitation, RNA pull-down, and minigene-MS2 mutation experiments all confirm SRSF1 binding to the SRA1 exon 3 sequence. SRA1-L overexpression increases CD44 expression and promotes HCC cell migration/invasion in vitro and lung metastasis in vivo; SRA1-S (lacking exon 3) reverses these effects. RNA immunoprecipitation (RIP), cross-link immunoprecipitation (CLIP), RNA pull-down, minigene-MS2 mutation, siRNA/overexpression, in vivo lung metastasis model Cell death discovery High 34011971

Source papers

Stage 0 corpus · 130 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2002 Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. Proceedings of the National Academy of Sciences of the United States of America 1479 12477932
2002 The transcription factor Sox9 has essential roles in successive steps of the chondrocyte differentiation pathway and is required for expression of Sox5 and Sox6. Genes & development 1408 12414734
1994 Campomelic dysplasia and autosomal sex reversal caused by mutations in an SRY-related gene. Nature 1268 7990924
1994 Autosomal sex reversal and campomelic dysplasia are caused by mutations in and around the SRY-related gene SOX9. Cell 1239 8001137
2009 A census of human transcription factors: function, expression and evolution. Nature reviews. Genetics 1191 19274049
2007 The SRA protein Np95 mediates epigenetic inheritance by recruiting Dnmt1 to methylated DNA. Nature 985 17994007
2012 Genome-wide meta-analysis identifies 56 bone mineral density loci and reveals 14 loci associated with risk of fracture. Nature genetics 958 22504420
2017 Impact of cytosine methylation on DNA binding specificities of human transcription factors. Science (New York, N.Y.) 934 28473536
2018 VIRMA mediates preferential m6A mRNA methylation in 3'UTR and near stop codon and associates with alternative polyadenylation. Cell discovery 829 29507755
2012 Slug and Sox9 cooperatively determine the mammary stem cell state. Cell 815 22385965
2011 Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Briefings in bioinformatics 656 21873635
1999 A steroid receptor coactivator, SRA, functions as an RNA and is present in an SRC-1 complex. Cell 640 10199399
2004 Interactions between Sox9 and beta-catenin control chondrocyte differentiation. Genes & development 637 15132997
1998 Direct interaction of SRY-related protein SOX9 and steroidogenic factor 1 regulates transcription of the human anti-Müllerian hormone gene. Molecular and cellular biology 482 9774680
2011 Genome-wide association study identifies loci influencing concentrations of liver enzymes in plasma. Nature genetics 456 22001757
2004 The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). Genome research 438 15489334
2004 SOX9 is an intestine crypt transcription factor, is regulated by the Wnt pathway, and represses the CDX2 and MUC2 genes. The Journal of cell biology 406 15240568
2008 Recognition of hemi-methylated DNA by the SRA protein UHRF1 by a base-flipping mechanism. Nature 395 18772891
2008 Structural basis for recognition of hemi-methylated DNA by the SRA domain of human UHRF1. Nature 379 18772889
2008 The SRA domain of UHRF1 flips 5-methylcytosine out of the DNA helix. Nature 342 18772888
2004 Sra-1 and Nap1 link Rac to actin assembly driving lamellipodia formation. The EMBO journal 325 14765121
2009 Highly conserved non-coding elements on either side of SOX9 associated with Pierre Robin sequence. Nature genetics 320 19234473
2007 The SRA methyl-cytosine-binding domain links DNA and histone methylation. Current biology : CB 315 17239600
2003 Abi, Sra1, and Kette control the stability and localization of SCAR/WAVE to regulate the formation of actin-based protrusions. Current biology : CB 294 14588242
2004 Smad3 induces chondrogenesis through the activation of SOX9 via CREB-binding protein/p300 recruitment. The Journal of biological chemistry 278 15623506
2001 Sharp, an inducible cofactor that integrates nuclear receptor repression and activation. Genes & development 277 11331609
2017 SOX9 Is an Astrocyte-Specific Nuclear Marker in the Adult Brain Outside the Neurogenic Regions. The Journal of neuroscience : the official journal of the Society for Neuroscience 274 28336567
2016 SOX9 and the many facets of its regulation in the chondrocyte lineage. Connective tissue research 264 27128146
2004 ICBP90, an E2F-1 target, recruits HDAC1 and binds to methyl-CpG through its SRA domain. Oncogene 263 15361834
2014 SRA- and SET-domain-containing proteins link RNA polymerase V occupancy to DNA methylation. Nature 259 24463519
2006 The RNA helicases p68/p72 and the noncoding RNA SRA are coregulators of MyoD and skeletal muscle differentiation. Developmental cell 248 17011493
2000 Phosphorylation of SOX9 by cyclic AMP-dependent protein kinase A enhances SOX9's ability to transactivate a Col2a1 chondrocyte-specific enhancer. Molecular and cellular biology 233 10805756
2013 ERG induces androgen receptor-mediated regulation of SOX9 in prostate cancer. The Journal of clinical investigation 229 23426182
2001 A subfamily of RNA-binding DEAD-box proteins acts as an estrogen receptor alpha coactivator through the N-terminal activation domain (AF-1) with an RNA coactivator, SRA. The EMBO journal 229 11250900
2014 Hippo coactivator YAP1 upregulates SOX9 and endows esophageal cancer cells with stem-like properties. Cancer research 227 24906622
1998 Toward understanding SOX9 function in chondrocyte differentiation. Matrix biology : journal of the International Society for Matrix Biology 212 9569122
2008 Loss of SR-A and CD36 activity reduces atherosclerotic lesion complexity without abrogating foam cell formation in hyperlipidemic mice. Arteriosclerosis, thrombosis, and vascular biology 211 18948635
2011 Toward an understanding of the protein interaction network of the human liver. Molecular systems biology 207 21988832
2005 CRMP-2 is involved in kinesin-1-dependent transport of the Sra-1/WAVE1 complex and axon formation. Molecular and cellular biology 207 16260607
2010 Mediation of CTCF transcriptional insulation by DEAD-box RNA-binding protein p68 and steroid receptor RNA activator SRA. Genes & development 194 20966046
2013 A meta-analysis of thyroid-related traits reveals novel loci and gender-specific differences in the regulation of thyroid function. PLoS genetics 190 23408906
2012 Oncogenicity of the developmental transcription factor Sox9. Cancer research 188 22246670
1996 Sex reversal by loss of the C-terminal transactivation domain of human SOX9. Nature genetics 181 8640233
2003 Transcriptional co-activators CREB-binding protein and p300 regulate chondrocyte-specific gene expression via association with Sox9. The Journal of biological chemistry 180 12732631
2010 Multiple roles for the non-coding RNA SRA in regulation of adipogenesis and insulin sensitivity. PloS one 176 21152033
2005 Prostaglandin D2 induces nuclear import of the sex-determining factor SOX9 via its cAMP-PKA phosphorylation. The EMBO journal 174 15889150
2001 Identification of human-infective trypanosomes in animal reservoir of sleeping sickness in Uganda by means of serum-resistance-associated (SRA) gene. Lancet (London, England) 161 11755607
1993 Assignment of an autosomal sex reversal locus (SRA1) and campomelic dysplasia (CMPD1) to 17q24.3-q25.1. Nature genetics 158 8348155
2011 Recognition of 5-hydroxymethylcytosine by the Uhrf1 SRA domain. PloS one 145 21731699
2008 Three SRA-domain methylcytosine-binding proteins cooperate to maintain global CpG methylation and epigenetic silencing in Arabidopsis. PLoS genetics 134 18704160
2008 SRA-domain proteins required for DRM2-mediated de novo DNA methylation. PLoS genetics 128 19043555
1984 Mutagenesis of the region between env and src of the SR-A strain of Rous sarcoma virus for the purpose of constructing helper-independent vectors. Virology 126 6330999
2010 Steroid receptor RNA activator protein binds to and counteracts SRA RNA-mediated activation of MyoD and muscle differentiation. Nucleic acids research 117 20855289
1999 Analysis of macrophage scavenger receptor (SR-A) expression in human aortic atherosclerotic lesions. Arteriosclerosis, thrombosis, and vascular biology 117 10073945
2020 Formononetin attenuates atherosclerosis via regulating interaction between KLF4 and SRA in apoE-/- mice. Theranostics 112 31938053
2013 Hypoxia enhances lipid uptake in macrophages: role of the scavenger receptors Lox1, SRA, and CD36. Atherosclerosis 112 23706521
2012 The SRA protein UHRF1 promotes epigenetic crosstalks and is involved in prostate cancer progression. Oncogene 112 22330138
2006 SLIRP, a small SRA binding protein, is a nuclear receptor corepressor. Molecular cell 109 16762838
2012 Internalization of modified lipids by CD36 and SR-A leads to hepatic inflammation and lysosomal cholesterol storage in Kupffer cells. PloS one 106 22470565
2010 Silencing of either SR-A or CD36 reduces atherosclerosis in hyperlipidaemic mice and reveals reciprocal upregulation of these receptors. Cardiovascular research 102 20634212
2014 SRA gene knockout protects against diet-induced obesity and improves glucose tolerance. The Journal of biological chemistry 101 24675075
2016 The lncRNA SLNCR1 Mediates Melanoma Invasion through a Conserved SRA1-like Region. Cell reports 100 27210747
2007 Steroid receptor RNA activator (SRA1): unusual bifaceted gene products with suspected relevance to breast cancer. Nuclear receptor signaling 100 17710122
2016 LncRNA SRA promotes hepatic steatosis through repressing the expression of adipose triglyceride lipase (ATGL). Scientific reports 96 27759039
2004 Interchangeable functions of Arabidopsis PIROGI and the human WAVE complex subunit SRA1 during leaf epidermal development. Development (Cambridge, England) 96 15294869
2011 A dual flip-out mechanism for 5mC recognition by the Arabidopsis SUVH5 SRA domain and its impact on DNA methylation and H3K9 dimethylation in vivo. Genes & development 93 21245167
2009 Dax-1 and steroid receptor RNA activator (SRA) function as transcriptional coactivators for steroidogenic factor 1 in steroidogenesis. Molecular and cellular biology 91 19188450
2014 Structural basis for hydroxymethylcytosine recognition by the SRA domain of UHRF2. Molecular cell 90 24813944
1994 The serum resistance-associated (SRA) gene of Trypanosoma brucei rhodesiense encodes a variant surface glycoprotein-like protein. Molecular and biochemical parasitology 90 7739673
2013 RNA-induced silencing complex (RISC) Proteins PACT, TRBP, and Dicer are SRA binding nuclear receptor coregulators. Proceedings of the National Academy of Sciences of the United States of America 87 23550157
2006 Alternative splicing of the first intron of the steroid receptor RNA activator (SRA) participates in the generation of coding and noncoding RNA isoforms in breast cancer cell lines. DNA and cell biology 87 16848684
2010 Protein complexes containing CYFIP/Sra/PIR121 coordinate Arf1 and Rac1 signalling during clathrin-AP-1-coated carrier biogenesis at the TGN. Nature cell biology 86 20228810
1998 A naturally occurring isoform of the human macrophage scavenger receptor (SR-A) gene generated by alternative splicing blocks modified LDL uptake. Journal of lipid research 80 9548586
2008 Structure and hemimethylated CpG binding of the SRA domain from human UHRF1. The Journal of biological chemistry 79 18945682
2008 HBEGF, SRA1, and IK: Three cosegregating genes as determinants of cardiomyopathy. Genome research 79 19064678
2010 Clinical and immunological determinants of severe/refractory asthma (SRA): association with Staphylococcal superantigen-specific IgE antibodies. Allergy 78 20973803
2009 Increasing the relative expression of endogenous non-coding Steroid Receptor RNA Activator (SRA) in human breast cancer cells using modified oligonucleotides. Nucleic acids research 77 19483093
2009 Pattern recognition scavenger receptor SRA/CD204 down-regulates Toll-like receptor 4 signaling-dependent CD8 T-cell activation. Blood 69 19349620
2010 EGb761 ameliorates the formation of foam cells by regulating the expression of SR-A and ABCA1: role of haem oxygenase-1. Cardiovascular research 66 20615914
2018 Expression Analysis of MALAT1, GAS5, SRA, and NEAT1 lncRNAs in Breast Cancer Tissues from Young Women and Women over 45 Years of Age. Molecular therapy. Nucleic acids 65 30126830
2012 PKCδ signalling regulates SR-A and CD36 expression and foam cell formation. Cardiovascular research 63 22687273
2005 Group V secretory phospholipase A2-modified low density lipoprotein promotes foam cell formation by a SR-A- and CD36-independent process that involves cellular proteoglycans. The Journal of biological chemistry 63 16040605
2001 Escherichia coli ribosome-associated protein SRA, whose copy number increases during stationary phase. Journal of bacteriology 63 11292794
2005 Efficacy and tolerability of darifenacin, a muscarinic M3 selective receptor antagonist (M3 SRA), compared with oxybutynin in the treatment of patients with overactive bladder. World journal of urology 62 16096831
2005 Defective microarchitecture of the spleen marginal zone and impaired response to a thymus-independent type 2 antigen in mice lacking scavenger receptors MARCO and SR-A. Journal of immunology (Baltimore, Md. : 1950) 62 16339556
2020 Xenbase: deep integration of GEO & SRA RNA-seq and ChIP-seq data in a model organism database. Nucleic acids research 59 31733057
2006 The calcineurin regulator sra plays an essential role in female meiosis in Drosophila. Current biology : CB 59 16860743
2010 Research resource: expression profiling reveals unexpected targets and functions of the human steroid receptor RNA activator (SRA) gene. Molecular endocrinology (Baltimore, Md.) 57 20219889
2019 Structure Relaxation Approximation (SRA) for Elucidation of Protein Structures from Ion Mobility Measurements. The journal of physical chemistry. B 53 30866623
2003 Ligand-independent coactivation of ERalpha AF-1 by steroid receptor RNA activator (SRA) via MAPK activation. The Journal of steroid biochemistry and molecular biology 53 12943696
2013 RNA helicase Ddx5 and the noncoding RNA SRA act as coactivators in the Notch signaling pathway. Biochimica et biophysica acta 52 23396200
2002 Synergically increased expression of CD36, CLA-1 and CD68, but not of SR-A and LOX-1, with the progression to foam cells from macrophages. Journal of atherosclerosis and thrombosis 52 12238639
1994 A point mutation leads to an unpaired cysteine residue and a molecular weight polymorphism of a functional platelet beta 3 integrin subunit. The Sra alloantigen system of GPIIIa. The Journal of biological chemistry 52 8132570
2018 Up-Regulation of Long Noncoding RNA SRA Promotes Cell Growth, Inhibits Cell Apoptosis, and Induces Secretion of Estradiol and Progesterone in Ovarian Granular Cells of Mice. Medical science monitor : international medical journal of experimental and clinical research 48 29674607
1992 Only the serum-resistant bloodstream forms of Trypanosoma brucei rhodesiense express the serum resistance associated (SRA) protein. Annales de la Societe belge de medecine tropicale 48 1417165
2022 Dextran sulfate-based MMP-2 enzyme-sensitive SR-A receptor targeting nanomicelles for the treatment of rheumatoid arthritis. Drug delivery 47 35119317
2004 Sra-1 interacts with Kette and Wasp and is required for neuronal and bristle development in Drosophila. Development (Cambridge, England) 47 15269173
2003 Activation of signaling pathways by putative scavenger receptor class A (SR-A) ligands requires CD14 but not SR-A. Biochemical and biophysical research communications 46 14521944
2001 Cell surface regulation of silica-induced apoptosis by the SR-A scavenger receptor in a murine lung macrophage cell line (MH-S). Toxicology and applied pharmacology 45 11437644
2018 New Insights Into the Long Non-coding RNA SRA: Physiological Functions and Mechanisms of Action. Frontiers in medicine 44 30238005
2011 Calcineurin and its regulator sra/DSCR1 are essential for sleep in Drosophila. The Journal of neuroscience : the official journal of the Society for Neuroscience 44 21900555
2010 Dax1 up-regulates Oct4 expression in mouse embryonic stem cells via LRH-1 and SRA. Molecular endocrinology (Baltimore, Md.) 44 20943815
2011 Targeting the immunoregulator SRA/CD204 potentiates specific dendritic cell vaccine-induced T-cell response and antitumor immunity. Cancer research 43 21914786
2010 Calcineurin and its regulation by Sra/RCAN is required for completion of meiosis in Drosophila. Developmental biology 43 20561515
2016 Tandem virtual screening targeting the SRA domain of UHRF1 identifies a novel chemical tool modulating DNA methylation. European journal of medicinal chemistry 42 27049577
2013 SR-A and SREC-I are Kupffer and endothelial cell receptors for helper-dependent adenoviral vectors. Molecular therapy : the journal of the American Society of Gene Therapy 39 23358188
2015 Association of the Long Non-coding RNA Steroid Receptor RNA Activator (SRA) with TrxG and PRC2 Complexes. PLoS genetics 36 26496121
2007 The DEAD-box p68/p72 proteins and the noncoding RNA steroid receptor activator SRA: eclectic regulators of disparate biological functions. Cell cycle (Georgetown, Tex.) 36 17495528
2009 SRA and its binding partners: an expanding role for RNA-binding coregulators in nuclear receptor-mediated gene regulation. Critical reviews in biochemistry and molecular biology 35 19280430
2008 SR-A, MARCO and TLRs differentially recognise selected surface proteins from Neisseria meningitidis: an example of fine specificity in microbial ligand recognition by innate immune receptors. Journal of innate immunity 35 20375573
2003 The C. elegans G-protein-coupled receptor SRA-13 inhibits RAS/MAPK signalling during olfaction and vulval development. Development (Cambridge, England) 35 12736202
2005 Pathological roles of advanced glycation end product receptors SR-A and CD36. Annals of the New York Academy of Sciences 34 16037291
2022 Using GenBank and SRA. Methods in molecular biology (Clifton, N.J.) 33 35037198
2016 Silencing of SRA1 Regulates ER Expression and Attenuates the Growth of Stromal Cells in Ovarian Endometriosis. Reproductive sciences (Thousand Oaks, Calif.) 33 27694140
2018 Targeting the SET and RING-associated (SRA) domain of ubiquitin-like, PHD and ring finger-containing 1 (UHRF1) for anti-cancer drug development. Oncotarget 32 29899856
2010 The scavenger receptors SRA-1 and SREC-I cooperate with TLR2 in the recognition of the hepatitis C virus non-structural protein 3 by dendritic cells. Journal of hepatology 32 20338659
2017 Roles of the Exogenous H2S-Mediated SR-A Signaling Pathway in Renal Ischemia/ Reperfusion Injury in Regulating Endoplasmic Reticulum Stress-Induced Autophagy in a Rat Model. Cellular physiology and biochemistry : international journal of experimental cellular physiology, biochemistry, and pharmacology 30 28472786
2014 Protective or deleterious role of scavenger receptors SR-A and CD36 on host resistance to Staphylococcus aureus depends on the site of infection. PloS one 29 24498223
2019 LncRNA-SRA1 Suppresses Osteosarcoma Cell Proliferation While Promoting Cell Apoptosis. Technology in cancer research & treatment 28 31106680
2016 Genetic variants in lncRNA SRA and risk of breast cancer. Oncotarget 28 26967566
2011 Recruitment of Dnmt1 roles of the SRA protein Np95 (Uhrf1) and other factors. Progress in molecular biology and translational science 28 21507355
2021 SRSF1 promotes the inclusion of exon 3 of SRA1 and the invasion of hepatocellular carcinoma cells by interacting with exon 3 of SRA1pre-mRNA. Cell death discovery 26 34011971
2008 The RNA coregulator SRA, its binding proteins and nuclear receptor signaling activity. IUBMB life 26 18380007
2004 Human infectivity trait in Trypanosoma brucei: stability, heritability and relationship to sra expression. Parasitology 26 15521633
2012 SR-A deficiency reduces myocardial ischemia/reperfusion injury; involvement of increased microRNA-125b expression in macrophages. Biochimica et biophysica acta 25 23123599
2011 SR-A ligand and M-CSF dynamically regulate SR-A expression and function in primary macrophages via p38 MAPK activation. BMC immunology 25 21736734
2011 Suppression of antigen-specific CD4+ T cell activation by SRA/CD204 through reducing the immunostimulatory capability of antigen-presenting cell. Journal of molecular medicine (Berlin, Germany) 25 22083206
2015 Recognition and cleavage of 5-methylcytosine DNA by bacterial SRA-HNH proteins. Nucleic acids research 24 25564526
2014 Structure and function of steroid receptor RNA activator protein, the proposed partner of SRA noncoding RNA. Journal of molecular biology 23 24486609
2011 Inflammatory stress exacerbates lipid-mediated renal injury in ApoE/CD36/SRA triple knockout mice. American journal of physiology. Renal physiology 23 21795641