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Showing TAFA4FAM19A4 is a alias.

TAFA4

Chemokine-like protein TAFA-4 · UniProt Q96LR4

Length
140 aa
Mass
15.7 kDa
Annotated
2026-06-10
34 papers in source corpus 11 papers cited in narrative 11 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 7/7 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

TAFA4 (FAM19A4) is a secreted neurokine produced by C-low-threshold mechanoreceptors (C-LTMRs), a subset of GINIP+ sensory neurons, that links somatosensory neurons to neuronal excitability control and innate immune regulation (PMID:24139797, PMID:34012116). As a marker of C-LTMRs, neuron-derived TAFA4 exerts an analgesic role: TAFA4-null mice show enhanced mechanical and chemical hypersensitivity with hyperexcitable spinal lamina II interneurons, and recombinant TAFA4 reverses this excitability and carrageenan-induced hypersensitivity (PMID:24139797). This antihypersensitive action operates through low-density lipoprotein receptor-related proteins (LRPs), with TAFA4 normalizing injury-induced changes in A-type K+ current and Ih in spinal interneurons in an LRP-dependent manner (PMID:34706225). Independently, TAFA4 functions as a ligand for formyl peptide receptor 1 (FPR1), driving macrophage chemotaxis, phagocytosis, Akt phosphorylation, and ROS release (PMID:25109685). Across tissues TAFA4 reprograms innate immune cells toward anti-inflammatory, pro-repair states, promoting macrophage M2 polarization and IL-10 production—via an LRP1-dependent route in dorsal root ganglia—to limit fibrosis, neuropathic pain, NLRP3 inflammasome activation, and post-infection inflammation (PMID:34012116, PMID:40381955, PMID:41666872, PMID:41747735). In adaptive and allergic immunity, TAFA4 signals through FPR1-MyD88-AKT in dendritic cells to induce Tr1 cells and through a PTEN-PU.1 axis in mast cells to restrict FcεRI expression, attenuating allergic rhinitis (PMID:36316414, PMID:36368013). In contrast, during sepsis TAFA4 enhances neutrophil ROS and NET formation through p38 MAPK, where its deficiency improves survival (PMID:41517951).

Mechanistic history

Synthesis pass · year-by-year structured walk · 9 steps
  1. 2013 High

    Established that a C-LTMR-specific secreted factor controls somatosensory thresholds, defining TAFA4 as an endogenous analgesic neuromodulator rather than a passive neuronal marker.

    Evidence Genetic labeling, TAFA4-null mice, spinal electrophysiology, and rescue by intrathecal/bath recombinant TAFA4 across multiple pain models

    PMID:24139797

    Open questions at the time
    • Receptor mediating the neuronal excitability effect not identified in this study
    • Molecular target of the excitability change (ion channel) not resolved
  2. 2014 High

    Identified a receptor for TAFA4, showing it acts as an FPR1 ligand to control macrophage chemotaxis and effector functions, opening an immune dimension to a neuronal factor.

    Evidence Radioligand binding, receptor internalization and blockade assays, in vitro chemotaxis, in vivo phagocytosis, and Akt phosphorylation western blot

    PMID:25109685

    Open questions at the time
    • Whether FPR1 mediates the in vivo analgesic effect not tested
    • No structural basis for TAFA4-FPR1 binding
  3. 2021 High

    Connected neuron-derived TAFA4 to tissue repair via a defined TAFA4-IL-10 macrophage axis, establishing a neuroimmune circuit controlling dermal macrophage survival and inflammatory tone.

    Evidence Conditional sensory neuron ablation, Tafa4-deficient mice, UV-skin damage model, in vitro macrophage stimulation, flow cytometry, histology

    PMID:34012116

    Open questions at the time
    • Receptor on dermal macrophages not defined in this study
    • Direct vs indirect induction of IL-10 not fully separated
  4. 2021 High

    Resolved the receptor requirement for TAFA4's analgesic action, showing LRPs mediate normalization of specific spinal interneuron ion currents.

    Evidence Multiple pain models, RAP pharmacological blockade, and electrophysiological recording of A-type K+ current and Ih in spinal interneurons

    PMID:34706225

    Open questions at the time
    • Specific LRP family member not pinpointed
    • Downstream signaling from LRP to ion channels unresolved
  5. 2022 Medium

    Extended TAFA4 signaling to dendritic cells, defining an FPR1-MyD88-AKT pathway that drives regulatory T cell induction and dampens allergic responses.

    Evidence Mouse allergic rhinitis model, in vitro DC stimulation, pathway inhibitor assays, flow cytometry for Tr1 cells

    PMID:36316414

    Open questions at the time
    • Pathway assigned by inhibitors without mutagenesis
    • Single lab
  6. 2023 Medium

    Identified a distinct mast-cell mechanism, the TAFA4-PTEN-PU.1 axis restricting Fcer1g transcription, broadening TAFA4's suppression of allergic activation.

    Evidence Patient nasal secretion ELISA, mouse allergic rhinitis model, in vitro mast cell stimulation, Fcer1g transcriptional analysis

    PMID:36368013

    Open questions at the time
    • No direct binding or receptor identified for the mast cell effect
    • Mechanistic link from receptor to PTEN unestablished
  7. 2025 Medium

    Demonstrated that boosting TAFA4 alleviates neuropathic pain via LRP1-dependent macrophage M2 polarization, unifying analgesic and immunomodulatory functions in a gain-of-function setting.

    Evidence CCI rat model with scAAV-mediated TAFA4 overexpression, flow cytometry, western blot, immunofluorescence, ELISA

    PMID:40381955

    Open questions at the time
    • Mechanistic detail limited
    • Relationship between LRP1 macrophage signaling and the earlier neuronal LRP effect not reconciled
  8. 2026 Medium

    Showed a context-dependent pro-inflammatory role in sepsis, where TAFA4 enhances neutrophil ROS and NET formation via p38 MAPK and its loss is protective, revealing the factor is not uniformly anti-inflammatory.

    Evidence Fam19a4 knockout CLP sepsis model, bulk RNA-seq, p38 inhibitor SB203580, ROS bioluminescence tracking, in vitro neutrophil assays

    PMID:41517951

    Open questions at the time
    • Receptor mediating neutrophil effect not identified
    • Source of TAFA4 in sepsis not defined
  9. 2026 Medium

    Generalized the TAFA4-IL-10 anti-inflammatory axis to disc degeneration and viral infection, establishing TAFA4 as a neuron-to-macrophage signal resolving inflammation across diverse pathologies.

    Evidence IVDD model with lentiviral TAFA4 knockdown and neuron-macrophage co-culture; HSV-1 DRG infection model with genetic/pharmacological dissection of TAFA4 vs substance P

    PMID:41666872 PMID:41747735

    Open questions at the time
    • Receptor mediating M2/IL-10 induction not resolved in these systems
    • NLRP3 suppression mechanism beyond ROS not defined

Open questions

Synthesis pass · forward-looking unresolved questions
  • How a single secreted factor selects among distinct receptors (FPR1 vs LRP/LRP1) and opposing immune outcomes (anti-inflammatory IL-10/M2 vs pro-inflammatory neutrophil ROS) in a cell- and context-specific manner remains unresolved.
  • No structural or biochemical basis distinguishing FPR1 vs LRP engagement
  • Determinants of pro- vs anti-inflammatory switching unknown
  • Receptor mediating spinal neuronal excitability changes not molecularly defined

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0098772 molecular function regulator activity 3 GO:0048018 receptor ligand activity 2
Localization
GO:0005576 extracellular region 3
Pathway
R-HSA-168256 Immune System 5 R-HSA-162582 Signal Transduction 3 R-HSA-112316 Neuronal System 2
Partners

Evidence

Reading pass · 11 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2013 TAFA4 is a specific marker of C-low-threshold mechanoreceptors (C-LTMRs). TAFA4-null mice exhibit enhanced mechanical and chemical hypersensitivity following inflammation and nerve injury, with increased excitability of spinal cord lamina IIi neurons. Intrathecal or bath application of recombinant TAFA4 protein reversed this increased excitability, and intrathecal administration strongly reversed carrageenan-induced mechanical hypersensitivity in wild-type mice, establishing an analgesic role for C-LTMR-derived TAFA4 in modulating neuronal excitability and somatic sensation thresholds. Genetic labeling, electrophysiological recordings, TAFA4-null mouse model (loss-of-function), intrathecal and bath application of recombinant TAFA4 protein Cell reports High 24139797
2014 FAM19A4 (TAFA4) is a ligand of formyl peptide receptor 1 (FPR1). The mature protein is composed of 95 amino acids. FAM19A4 shows chemotactic activity on macrophages, enhances macrophage phagocytosis of zymosan (with increased Akt phosphorylation), and increases reactive oxygen species (ROS) release. These activities were demonstrated by receptor internalization assays, radioligand binding assays, and receptor blockage experiments. Receptor internalization assay, radioligand binding assay, receptor blockage, in vitro chemotaxis assay, in vivo phagocytosis assay, western blot for Akt phosphorylation Cellular & molecular immunology High 25109685
2021 Sensory neuron-derived TAFA4, produced specifically by C-low threshold mechanoreceptors (a subset of GINIP+ neurons), promotes the tissue-repair function of dermal macrophages. In vivo, TAFA4-deficient mice showed defective tissue regeneration and dermal fibrosis after UV-induced skin damage. TAFA4 modulates the inflammatory profile of macrophages directly in vitro. A TAFA4-IL-10 axis was identified whereby TAFA4 promotes IL-10 production by dermal macrophages and ensures survival and maintenance of IL-10+TIM4+ dermal macrophages, reducing inflammation and promoting tissue regeneration. Conditional sensory neuron ablation, Tafa4-deficient mouse model, in vitro macrophage stimulation, in vivo UV-skin damage model, flow cytometry, histology Nature High 34012116
2021 TAFA4 reverses inflammatory, postoperative, and spared nerve injury (SNI)-induced mechanical hypersensitivity through functional low-density lipoprotein receptor-related proteins (LRPs), as their inhibition by RAP (receptor-associated protein) dose-dependently abolished TAFA4's antihypersensitive actions. SNI selectively decreases A-type K+ current (IA) in spinal lamina II outer excitatory interneurons and alters Ih in lamina II inner inhibitory interneurons; these ion current alterations were rescued by TAFA4 in an LRP-dependent manner. In vivo mechanical hypersensitivity assays, pharmacological inhibition (RAP), electrophysiological recordings of spinal interneuron ion currents, multiple pain models (inflammatory, postoperative, SNI) Cell reports High 34706225
2022 TAFA4 activates dendritic cells (DCs) in airway tissues, inducing IL-10 expression via the FPR1-MyD88-AKT signaling pathway. TAFA4 also promotes activities of c-Maf inducing protein in DCs. Co-administration of TAFA4 with allergen immunotherapy induced antigen-specific Tr1 regulatory T cells and attenuated allergic rhinitis response in mice. Mouse allergic rhinitis model, DC stimulation in vitro, signaling pathway inhibitor assays (FPR1-MyD88-AKT), flow cytometry for Tr1 cells NPJ vaccines Medium 36316414
2023 TAFA4 modulates FcεRI expression in mast cells through the TAFA4-PTEN-PU.1 signaling axis, which restricts transcription of Fcer1g (FcεRI γ gene). TAFA4 suppressed antigen-related mast cell activation and attenuated experimental allergic rhinitis. Negative correlation between TAFA4 and tryptase levels in nasal secretions from allergic rhinitis patients was observed. ELISA (nasal secretions from patients), mouse allergic rhinitis model, in vitro mast cell stimulation, transcriptional analysis of Fcer1g, PTEN-PU.1 pathway interrogation Clinical and experimental immunology Medium 36368013
2025 TAFA4 overexpression in dorsal root ganglia (DRG) ameliorates neuropathic pain by promoting macrophage M2 polarization in the DRGs. Mechanistically, TAFA4 modulates macrophage function in a lipoprotein receptor-related protein 1 (LRP1)-dependent manner. TAFA4 overexpression (via scAAV) increased IL-10 concentrations in DRG and inhibited pro-inflammatory mediators after chronic constriction injury. Chronic constriction injury (CCI) rat model, scAAV-mediated TAFA4 overexpression, flow cytometry, western blot, immunofluorescence, ELISA Neurochemistry international Medium 40381955
2026 FAM19A4 enhances neutrophil reactive oxygen species (ROS) production specifically through p38 MAPK signaling activation. FAM19A4 deficiency (Fam19a4 knockout mice) in a cecal ligation and puncture (CLP) sepsis model improved survival and reduced multiorgan injury, reduced neutrophil and macrophage counts in lungs and liver, and decreased neutrophil extracellular trap (NET) formation. In vitro, FAM19A4 enhanced neutrophil phagocytosis and ROS generation but did not affect LPS-induced chemotaxis. Fam19a4 knockout mouse CLP sepsis model, bulk RNA sequencing, western blot, p38 MAPK inhibitor (SB203580) treatment, bioluminescence ROS tracking, flow cytometry, in vitro neutrophil assays Acta biochimica et biophysica Sinica Medium 41517951
2026 TAFA4 acts as a neuron-derived mediator in intervertebral disc degeneration (IVDD). GINIP+ sensory neurons secrete TAFA4 in degenerative discs. In vitro, GINIP+ neurons promoted macrophage M2 polarization and IL-10 production while suppressing TNF-α and IL-1β; these effects were reversed by TAFA4 knockdown. TAFA4 attenuated ROS-dependent NLRP3 inflammasome activation in macrophages. In vivo IVDD model with lentiviral TAFA4 knockdown, in vitro neuron-macrophage co-culture, flow cytometry, immunofluorescence, ELISA, western blot Neurospine Medium 41666872
2026 During herpes simplex virus type 1 infection, sensory neurons produce TAFA4. In infected dorsal root ganglia (DRGs), a TAFA4-IL-10 pathway promotes the resolution of inflammation after viral clearance, functioning independently from substance P actions in skin. This neuroimmune regulatory pathway reduces detrimental impact of infection on host fitness without directly altering pathogen elimination. HSV-1 mouse infection model, tissue-specific analysis of DRG vs. skin, genetic and pharmacological dissection of SP and TAFA4 pathways Immunity Medium 41747735
2024 In a Shank3 mouse model of autism, C-fiber low-threshold mechanoreceptors (C-LTMRs) are hyporesponsive and TAFA4 is transcriptomically downregulated. TAFA4 injection reduced spontaneous scratching response to skin deformation but failed to restore itch sensitivity, suggesting TAFA4 modulates one component of mechanical itch but not the full alloknesis pathway. Ex vivo electrophysiology of C-LTMRs, transcriptomic analysis, TAFA4 pharmacological injection, Shank3 ΔC/ΔC mouse model bioRxivpreprint Low bio_10.1101_2024.12.29.630575

Source papers

Stage 0 corpus · 34 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2021 Sensory neuron-derived TAFA4 promotes macrophage tissue repair functions. Nature 157 34012116
2014 Methylation analysis of the FAM19A4 gene in cervical scrapes is highly efficient in detecting cervical carcinomas and advanced CIN2/3 lesions. Cancer prevention research (Philadelphia, Pa.) 115 25281488
2013 TAFA4, a chemokine-like protein, modulates injury-induced mechanical and chemical pain hypersensitivity in mice. Cell reports 114 24139797
2020 Methylation markers FAM19A4 and miR124-2 as triage strategy for primary human papillomavirus screen positive women: A large European multicenter study. International journal of cancer 104 32997803
2016 Validation of the FAM19A4/mir124-2 DNA methylation test for both lavage- and brush-based self-samples to detect cervical (pre)cancer in HPV-positive women. Gynecologic oncology 90 26921784
2018 Cervical cancer risk in HPV-positive women after a negative FAM19A4/mir124-2 methylation test: A post hoc analysis in the POBASCAM trial with 14 year follow-up. International journal of cancer 84 29663363
2014 FAM19A4 is a novel cytokine ligand of formyl peptide receptor 1 (FPR1) and is able to promote the migration and phagocytosis of macrophages. Cellular & molecular immunology 71 25109685
2022 Clinical Regression of High-Grade Cervical Intraepithelial Neoplasia Is Associated With Absence of FAM19A4/miR124-2 DNA Methylation (CONCERVE Study). Journal of clinical oncology : official journal of the American Society of Clinical Oncology 66 35512257
2019 FAM19A4/miR124-2 methylation in invasive cervical cancer: A retrospective cross-sectional worldwide study. International journal of cancer 65 31390052
2015 Comparing the performance of FAM19A4 methylation analysis, cytology and HPV16/18 genotyping for the detection of cervical (pre)cancer in high-risk HPV-positive women of a gynecologic outpatient population (COMETH study). International journal of cancer 62 26317579
2020 FAM19A4/miR124-2 methylation analysis as a triage test for HPV-positive women: cross-sectional and longitudinal data from a Dutch screening cohort. Clinical microbiology and infection : the official publication of the European Society of Clinical Microbiology and Infectious Diseases 47 32222459
2019 Long-term CIN3+ risk of HPV positive women after triage with FAM19A4/miR124-2 methylation analysis. Gynecologic oncology 43 31182225
2021 Classification of high-grade cervical intraepithelial neoplasia by p16ink4a , Ki-67, HPV E4 and FAM19A4/miR124-2 methylation status demonstrates considerable heterogeneity with potential consequences for management. International journal of cancer 39 33729551
2018 The clinical significance of FAM19A4 methylation in high-risk HPV-positive cervical samples for the detection of cervical (pre)cancer in Chinese women. BMC cancer 33 30486875
2019 Intra- and inter-laboratory agreement of the FAM19A4/mir124-2 methylation test: Results from an international study. Journal of clinical laboratory analysis 29 30758084
2022 Molecular Structure, Expression and Role of TAFA4 and its Receptor FPR1 in the Spinal Cord. Frontiers in cell and developmental biology 25 35712659
2021 TAFA4 relieves injury-induced mechanical hypersensitivity through LDL receptors and modulation of spinal A-type K+ current. Cell reports 23 34706225
2023 FAM19A4/miR124-2 Methylation Testing and Human Papillomavirus (HPV) 16/18 Genotyping in HPV-Positive Women Under the Age of 30 Years. Clinical infectious diseases : an official publication of the Infectious Diseases Society of America 21 35686306
2020 Expression of p16 and HPV E4 on biopsy samples and methylation of FAM19A4 and miR124-2 on cervical cytology samples in the classification of cervical squamous intraepithelial lesions. Cancer medicine 17 32022461
2024 FAM19A4 and hsa-miR124-2 Double Methylation as Screening for ASC-H- and CIN1 HPV-Positive Women. Pathogens (Basel, Switzerland) 16 38668267
2019 Role of FAM19A4/miR124-2 methylation analysis in predicting regression or non-regression of CIN2/3 lesions: a protocol of an observational longitudinal cohort study. BMJ open 14 31289088
2018 Defining hrHPV genotypes in cervical intraepithelial neoplasia by laser capture microdissection supports reflex triage of self-samples using HPV16/18 and FAM19A4/miR124-2 methylation. Gynecologic oncology 9 30219239
2023 Cervix cytology samples revealed increased methylation of the human markers FAM19A4/miR124-2 up to 8 years before adenocarcinoma. Acta obstetricia et gynecologica Scandinavica 7 37964497
2015 Efficient production of FAM19A4, a novel potential cytokine, in a stable optimized CHO-S cell system. Protein expression and purification 6 25979463
2022 TAFA4-IL-10 axis potentiate immunotherapy for airway allergy by induction of specific regulatory T cells. NPJ vaccines 5 36316414
2024 DNA methylation at individual CpG-sites of EPB41L3, HTERT and FAM19A4 are useful for detection of cervical high-grade squamous intraepithelial lesions (HSIL) or worse: Analysis of individual CpG-sites outperforms averaging. Tumour virus research 3 38960143
2023 Signals from the TAFA4-PTEN-PU.1 axis alleviate nasal allergy by modulating the expression of FcεRI in mast cells. Clinical and experimental immunology 3 36368013
2025 Overexpression of TAFA4 in the dorsal root ganglion ameliorates neuropathic pain in male rats through promoting macrophage M2-Skewing. Neurochemistry international 2 40381955
2026 Sensory neuron production of substance P and TAFA4 promotes disease tolerance during viral infection. Immunity 1 41747735
2025 FAM19A4 and miR124-2 methylation status in human papillomavirus-driven and human papillomavirus-negative oropharyngeal squamous cell carcinomas. Infectious agents and cancer 1 41094556
2017 Method for quantitative detection of FAM19A4 by flow cytometry using latex beads as solid carrier. Journal of bioscience and bioengineering 1 29167066
2026 FAM19A4 enhances neutrophil respiratory burst via p38 MAPK in lethal sepsis. Acta biochimica et biophysica Sinica 0 41517951
2026 TAFA4 Mitigates Intervertebral Disc Degeneration by Modulating Macrophage Polarization and Inhibiting ROS-NLRP3 Inflammasome Activation. Neurospine 0 41666872
2019 [Detection and analysis of FAM19A4 promoter methylation in cervical exfoliated cells]. Zhonghua yi xue za zhi 0 31269601

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