Affinage

CAVIN3

Caveolae-associated protein 3 · UniProt Q969G5

Length
261 aa
Mass
27.7 kDa
Annotated
2026-06-09
30 papers in source corpus 16 papers cited in narrative 16 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 7/7 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

CAVIN3 (cavin-3/SRBC/PRKCDBP) is a caveolae-associated adapter protein that governs caveolae dynamics and intracellular trafficking while coupling caveolar membrane organization to growth-factor signaling and stress responses (PMID:19262564, PMID:24069528, PMID:25588833). It localizes to caveolae through a leucine zipper required for co-precipitation with caveolin-1 and is recruited there by cavin1, whose trimeric N-terminus it binds in competition with cavin2 to define distinct cavin subcomplexes; CAVIN3 is enriched at deeply invaginated caveolae and shifts the population toward short-lived, dynamic structures, and its loss markedly impairs microtubule-based trafficking of caveolar vesicles (cavicles) (PMID:19262564, PMID:25588833). CAVIN3 also binds PKCδ as a member of the STICK superfamily (PMID:19262564). Through anchoring of caveolae to the membrane skeleton via myosin-1c, CAVIN3 sets the balance between ERK and Akt signaling: its loss suppresses ERK while elevating Akt (via reduced EGR1/PTEN), and CAVIN3 knockout mice show increased lactate production and cachexia (PMID:24069528). Beyond caveolae, CAVIN3 acts as a stress-responsive tumor suppressor: it is released from disassembling caveolae upon UV and mechanical stress to directly interact with BRCA1 and sustain the BRCA1 A-complex DNA-repair pathway, with CAVIN3-deficient cells showing PARP-inhibitor sensitivity reversed by 53BP1 co-depletion (PMID:34142659). It stabilizes p53 to drive p53-dependent G1 arrest and apoptosis (PMID:18059034), and its transcription is directly activated by NF-κB downstream of TNFα (PMID:21980136). CAVIN3 additionally functions in a caveolae-independent context as a cytoplasmic PER2-interacting protein that modulates circadian period length in a PKCδ-binding-site-dependent manner (PMID:23079727).

Mechanistic history

Synthesis pass · year-by-year structured walk · 15 steps
  1. 2008 Medium

    Established CAVIN3 as a growth-suppressive protein acting through p53, providing the first mechanistic link to tumor suppression.

    Evidence Stable/transient overexpression with p53 functional blockade and xenograft assays measuring p53 target gene induction

    PMID:18059034

    Open questions at the time
    • Did not define how CAVIN3 physically stabilizes p53
    • No endogenous loss-of-function tumor data
  2. 2009 High

    Identified the molecular basis for CAVIN3's caveolar localization and its requirement for caveolar vesicle trafficking, establishing it as a caveolae adapter.

    Evidence Reciprocal Co-IP with caveolin-1, leucine-zipper mutagenesis, PKCδ binding, and live-cell imaging of cavicle trafficking

    PMID:19262564

    Open questions at the time
    • Did not resolve how CAVIN3 controls cavicle motility on microtubules
    • Functional role of PKCδ binding left open
  3. 2011 High

    Showed CAVIN3 is a direct NF-κB transcriptional target induced by TNFα, linking its expression to inflammatory signaling and apoptotic sensitivity.

    Evidence Luciferase reporter, ChIP, κB-site mutagenesis, RelA transfection in tumor cells

    PMID:21980136

    Open questions at the time
    • Did not connect transcriptional induction to a downstream CAVIN3 effector mechanism
  4. 2012 Medium

    Revealed a caveolae-independent role: CAVIN3 binds PER2 in the cytoplasm to set circadian period length, expanding its functional repertoire.

    Evidence Co-IP of PER2, RNAi/overexpression circadian period assays, PKCδ-binding-site mutagenesis in fibroblasts

    PMID:23079727

    Open questions at the time
    • The kinase mediating the PKCδ-site-dependent effect remains unidentified
    • Direct vs indirect PER2 interaction not resolved
  5. 2013 High

    Defined CAVIN3 as a switch between ERK and Akt signaling by anchoring caveolae to the membrane skeleton through myosin-1c, connecting caveolar architecture to signaling outcomes and organismal metabolism.

    Evidence Knockout mice, gain/loss-of-function, Co-IP with myosin-1c, ERK/Akt phosphorylation and metabolic assays

    PMID:24069528

    Open questions at the time
    • Mechanism by which CAVIN3 suppresses EGR1/PTEN not detailed
    • Tissue specificity of the signaling balance unresolved
  6. 2014 Medium

    Clarified that CAVIN3 is dispensable for caveolae formation and metabolic homeostasis in adipose and other tissues, distinguishing it from core caveolae structural cavins.

    Evidence Knockout mice with EM caveolae counting and metabolic phenotyping

    PMID:25036884

    Open questions at the time
    • Negative finding does not exclude tissue-specific roles tested elsewhere
  7. 2014 Medium

    Extended CAVIN3's tumor-suppressive function to invasion control via an AKT/MMP-9 axis.

    Evidence Overexpression/knockdown with MMP-9 zymography, migration assays, AKT phosphorylation western blot

    PMID:25520561

    Open questions at the time
    • How CAVIN3 prevents AKT dephosphorylation mechanistically unclear
  8. 2015 High

    Mapped the structural logic of cavin subcomplex assembly, showing CAVIN3's N-terminus binds a cavin1 trimer competitively with cavin2 and biases caveolae toward dynamic short-lived states.

    Evidence Co-IP, truncation/domain mapping, live-cell TIRF imaging of caveolae dynamics, siRNA

    PMID:25588833

    Open questions at the time
    • Stoichiometry of mixed cavin1/2/3 complexes not fully resolved
  9. 2017 Medium

    Demonstrated a tissue-specific requirement for CAVIN3 in caveolae maintenance in smooth muscle and an effect on NO-dependent vascular relaxation.

    Evidence Knockout mice, EM caveolae quantification, vascular contraction/relaxation assays, cavin-1/sGC western blots

    PMID:28285351

    Open questions at the time
    • Why dependence is restricted to smooth muscle unexplained
  10. 2019 Medium

    Identified ROR1 as a CAVIN3 partner required for its proper localization and caveolae-dependent endocytosis feeding pro-survival AKT signaling.

    Evidence Co-IP with site mapping, siRNA, endocytosis assays, fractionation, AKT readouts in lung adenocarcinoma

    PMID:30894682

    Open questions at the time
    • Direct vs scaffold-mediated ROR1 interaction not distinguished
  11. 2020 Medium

    Showed CAVIN3 promotes adipogenesis by modulating TACE/ADAM17-mediated shedding of the inhibitor Pref-1.

    Evidence Stable knockdown/overexpression, qRT-PCR, western blot, TACE activity and adipogenesis assays

    PMID:32679831

    Open questions at the time
    • How CAVIN3 regulates TACE activity mechanistically undefined
  12. 2020 Medium

    Revealed CAVIN3 as a host factor hijacked by hPIV-2 V protein, which stabilizes CAVIN3 in lipid rafts to support viral assembly and budding.

    Evidence Co-IP with domain mapping, pulse-chase degradation, siRNA, viral growth assays, lipid raft fractionation

    PMID:32425917

    Open questions at the time
    • Cellular function co-opted by V protein binding not defined
  13. 2021 High

    Established CAVIN3 as a stress-released caveolar protein that directly engages BRCA1 to support the BRCA1 A-complex DNA-repair pathway, mechanistically unifying caveolae disassembly with genome maintenance.

    Evidence CRISPR KO, label-free proteomics, cell-free direct interaction assays, UV-apoptosis, PARP-inhibitor sensitivity, 53BP1/BRCA1 epistasis

    PMID:34142659

    Open questions at the time
    • Precise binding interface with BRCA1 not structurally defined
    • Trigger signal linking caveolae disassembly to nuclear BRCA1 unclear
  14. 2023 Medium

    Linked CAVIN3 to NLRP3-dependent pyroptosis and identified a lncRNA/miRNA ceRNA axis controlling its expression in NSCLC.

    Evidence Pyroptosis assays (ASC specks, CASP1/IL-1β/GSDMD), dual-luciferase, RNA immunoprecipitation

    PMID:36696967

    Open questions at the time
    • How CAVIN3 activates NLRP3 mechanistically unknown
  15. 2025 Medium

    Placed CAVIN3 in endothelial neovascularization control through an ERK/JAG1 axis regulated by hypoxic ZEB1.

    Evidence siRNA in endothelial cells, CNV/OIR mouse models, ERK/JAG1 assays, ZEB1 ChIP/reporter, pericyte-EC co-culture

    PMID:40337864

    Open questions at the time
    • Connection to caveolar function in endothelium not established

Open questions

Synthesis pass · forward-looking unresolved questions
  • How CAVIN3's distinct caveolar, signaling, stress/DNA-repair, and circadian activities are coordinated within a single cell, and the structural basis for its multiple direct interactions, remain unresolved.
  • No unified structural model for CAVIN3's separable interaction modes
  • Switch governing caveolar vs cytoplasmic vs nuclear pools undefined

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0060090 molecular adaptor activity 3 GO:0098772 molecular function regulator activity 2
Localization
GO:0005886 plasma membrane 2 GO:0031410 cytoplasmic vesicle 2 GO:0005829 cytosol 1
Pathway
R-HSA-162582 Signal Transduction 2 R-HSA-5357801 Programmed Cell Death 2 R-HSA-1852241 Organelle biogenesis and maintenance 1 R-HSA-73894 DNA Repair 1 R-HSA-9909396 Circadian clock 1
Complex memberships
caveolae cavin coat (cavin1-cavin3 subcomplex)

Evidence

Reading pass · 16 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2009 SRBC/cavin-3 (CAVIN3) co-immunoprecipitates with caveolin-1, and a leucine zipper in SRBC is essential for both co-precipitation with caveolin-1 and localization to caveolae. SRBC binds PKCδ as a member of the STICK superfamily. SRBC remains associated with caveolin when caveolae bud to form vesicles (cavicles) that travel on microtubules, and in the absence of SRBC, intracellular cavicle traffic is markedly impaired. Co-immunoprecipitation, leucine zipper mutagenesis, live-cell imaging of cavicle trafficking The EMBO journal High 19262564
2013 Cavin-3 dictates the balance between ERK and Akt signaling: loss of cavin-3 increases Akt signaling at the expense of ERK, while gain of cavin-3 increases ERK signaling at the expense of Akt. Cavin-3 facilitates ERK signaling by anchoring caveolae to the membrane skeleton via myosin-1c; loss of this linkage reduces caveolae abundance and separates the ERK activation module from signaling receptors. Loss of cavin-3 also promotes Akt signaling through suppression of EGR1 and PTEN. In vivo consequences of cavin-3 knockout include increased lactate production and cachexia. Cavin-3 knockout mice, in vitro gain/loss-of-function, co-immunoprecipitation with myosin-1c, signaling pathway analysis (ERK/Akt phosphorylation), metabolic assays eLife High 24069528
2012 CAVIN-3 is a cytoplasmic PER2-interacting protein that influences circadian clock properties. Loss- and gain-of-function of CAVIN-3 shortened and lengthened, respectively, the circadian period in fibroblasts, and affected PER:CRY protein abundance and interaction. CAVIN-3 required its PKCδ-binding site to exert its effect on period length. Depletion of PKCδ alone had little effect, suggesting involvement of yet-uncharacterized kinases. CAVIN-3 activity in circadian gene expression was independent of caveolae. Co-immunoprecipitation (PER2 interaction), RNAi knockdown, overexpression, circadian period assays in fibroblasts, PKCδ-binding site mutagenesis EMBO reports Medium 23079727
2015 Cavin3 is targeted to caveolae by cavin1, where it interacts with the scaffolding domain of caveolin1 and promotes caveolae dynamics. The N-terminal region of cavin3 binds a trimer of the cavin1 N-terminus in competition with a homologous cavin2 region, indicating that cavins form distinct subcomplexes through their N-terminal regions. Cavin3 is enriched at deeply invaginated caveolae, and loss of cavin3 increases the proportion of stable caveolae and decreases short-lived caveolae at the membrane. Co-immunoprecipitation, truncation/domain mapping, live-cell TIRF imaging of caveolae dynamics, siRNA knockdown Journal of cell science High 25588833
2008 hSRBC (CAVIN3) increases p53 protein stability and expression of p53 target genes (p21Waf1, PUMA, NOXA). hSRBC-mediated cell cycle arrest and apoptosis were abolished by blockade of p53 function. Stable expression of hSRBC led to G1 cell cycle arrest, apoptosis, suppressed colony forming ability and xenograft tumor growth, and elevated sensitivity to genotoxic agents. Stable and transient overexpression, p53 functional blockade, xenograft tumor assay, western blot for p53 target genes International journal of cancer Medium 18059034
2011 PRKCDBP (CAVIN3) gene transcription is directly activated by NF-κB in response to TNFα: luciferase reporter and ChIP assays demonstrate that TNFα-induced PRKCDBP promoter activity requires an intact κB site, and RelA transfection enhances PRKCDBP expression. PRKCDBP induction correlates with tumor cell sensitivity to TNFα-induced apoptosis. Luciferase reporter assay, chromatin immunoprecipitation (ChIP), RelA transfection, siRNA knockdown, xenograft assay Clinical cancer research High 21980136
2019 ROR1 binds CAVIN3 at a site distinct from those for CAV1 and CAVIN1; this interaction is required for proper CAVIN3 subcellular localization and caveolae-dependent endocytosis but not caveolae formation itself. The ROR1-CAVIN3 interaction facilitates caveolae trafficking linked to pro-survival AKT signaling from early endosomes in lung adenocarcinoma cells. Co-immunoprecipitation, domain mapping, siRNA knockdown, endocytosis assays, subcellular fractionation/localization, AKT signaling readouts Oncogene Medium 30894682
2021 Cavin3 is released from caveolae upon UV and mechanical stress-induced caveolae disassembly and directly interacts with BRCA1. Cavin3 deletion downregulates BRCA1 and BRCA1 A-complex components. Cavin3 sensitizes cancer cells to UV-induced apoptosis, and cavin3-deficient cells are sensitive to PARP inhibition; concomitant depletion of 53BP1 restored BRCA1-dependent sensitivity to PARP inhibition, placing cavin3 in the BRCA1 DNA repair pathway. Genome editing (CRISPR KO), label-free quantitative proteomics, cell-free expression interaction assays, RNAi depletion, UV-apoptosis assays, PARP inhibitor sensitivity, genetic epistasis (53BP1/BRCA1 double depletion) eLife High 34142659
2014 In cavin-3 knockout mice, loss of cavin-3 does not significantly affect caveolae abundance in adipose tissue or other widely studied tissues, and has no effect on body composition or glucose tolerance, indicating cavin-3 is not absolutely required for caveolae formation in these contexts. Cavin-3 knockout mouse generation, electron microscopy for caveolae counting, metabolic phenotyping (body weight, fat mass, glucose tolerance tests), microarray PloS one Medium 25036884
2017 Loss of cavin-3 in smooth muscle reduces cavin-1 protein levels by ~40% and reduces caveolae density by 40–45% in vascular and urinary bladder smooth muscle (but not in endothelial cells), demonstrating a tissue-specific role for cavin-3 in caveolae maintenance in smooth muscle. Enhanced nitric-oxide-dependent vascular relaxation was observed alongside elevated soluble guanylyl cyclase expression in KO mice. Cavin-3 knockout mice, electron microscopy (caveolae quantification), vascular contraction/relaxation assays, western blot for cavin-1/sGC protein levels Cell and tissue research Medium 28285351
2014 Cavin-3 overexpression in HT1080 fibrosarcoma cells reduces cell migration and inhibits PMA-induced MMP-9 secretion and gene expression, in part through prevention of AKT dephosphorylation. Cavin-3 gene silencing increases MMP-9 expression and secretion, establishing a cavin-3/AKT/MMP-9 signaling axis. Cavin-3 overexpression and siRNA knockdown, MMP-9 zymography/ELISA, cell migration assay, AKT phosphorylation western blot, PMA stimulation Cancer growth and metastasis Medium 25520561
2020 Cavin-3 knockdown impairs adipocyte differentiation; its overexpression accelerates adipogenesis. Mechanistically, Cavin-3 modulates TACE (ADAM17)-mediated shedding of Pref-1 (preadipocyte factor-1), a known inhibitor of adipogenesis. Cavin-3 silencing markedly increases Pref-1 during adipocyte maturation while decreasing expression of adipogenesis genes (PPARγ, FAS, aP2, Adipoq). Stable knockdown/overexpression cell lines, qRT-PCR, western blot, confocal immunofluorescence, TACE activity assay, adipogenesis assays International journal of molecular sciences Medium 32679831
2020 Human parainfluenza virus type 2 (hPIV-2) V protein binds directly to Cavin3 (N-terminal region of Cavin3 and Trp residues in C-terminal region of V protein are required), inhibits Cavin3 proteasomal degradation, increases Cavin3 abundance in lipid raft microdomains, and thereby facilitates viral assembly and budding. Cavin3 knockdown suppresses hPIV-2 growth without affecting viral entry, replication, transcription, or translation. Co-immunoprecipitation, pulse-chase proteasomal degradation assay, V protein overexpression, siRNA knockdown, viral growth assays, lipid raft fractionation Frontiers in microbiology Medium 32425917
2025 CAVIN3 deficiency in endothelial cells inhibits ERK phosphorylation, which downregulates Jagged 1 (JAG1) expression, thereby promoting vascular normalization in pathological neovascularization. Transcription factor ZEB1 regulates CAVIN3 transcription in endothelial cells under hypoxic conditions. CAVIN3 knockdown disrupts EC proliferation and vascular sprouting, restores pericyte-EC interactions. siRNA knockdown in endothelial cells, in vivo CNV and OIR mouse models, ERK phosphorylation western blot, JAG1 expression assays, ZEB1 ChIP/reporter assays, pericyte-EC co-culture JCI insight Medium 40337864
2024 A phosphomimic mutation in a Thr-Ser pair proximal to the disordered C-terminal half of the Cavin1 HR1 coiled-coil domain selectively abolishes Cavin2 and Cavin3 association with Cavin1, revealing that phosphorylation near this region regulates the formation of Cavin1-Cavin2 and Cavin1-Cavin3 subcomplexes. Nanobody development, X-ray crystal structure of nanobody-HR1 complex, phosphomimic mutagenesis, co-immunoprecipitation bioRxivpreprint Medium bio_10.1101_2024.11.26.625551
2023 PRKCDBP promotes pyroptosis by activating the NLRP3 inflammasome, resulting in cleavage of CASP1, IL-1β, and GSDMD in NSCLC cells. PRKCDBP is upregulated through a ceRNA mechanism in which lncRNA TCONS-14036 sequesters miR-1228-5p, relieving miR-1228-5p-mediated suppression of PRKCDBP. Flow cytometry, TUNEL assay, ASC speck formation, ELISA, dual-luciferase reporter assay, RNA immunoprecipitation, western blot, siRNA/overexpression Cell proliferation Medium 36696967

Source papers

Stage 0 corpus · 30 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2009 SRBC/cavin-3 is a caveolin adapter protein that regulates caveolae function. The EMBO journal 170 19262564
2013 Cavin-3 dictates the balance between ERK and Akt signaling. eLife 62 24069528
2011 Epigenetic alteration of PRKCDBP in colorectal cancers and its implication in tumor cell resistance to TNFα-induced apoptosis. Clinical cancer research : an official journal of the American Association for Cancer Research 48 21980136
2015 Cavin3 interacts with cavin1 and caveolin1 to increase surface dynamics of caveolae. Journal of cell science 45 25588833
2008 Frequent epigenetic inactivation of hSRBC in gastric cancer and its implication in attenuated p53 response to stresses. International journal of cancer 39 18059034
2005 Expression of the candidate tumor suppressor gene hSRBC is frequently lost in primary lung cancers with and without DNA methylation. Oncogene 39 15940253
2014 CRY1, CRY2 and PRKCDBP genetic variants in metabolic syndrome. Hypertension research : official journal of the Japanese Society of Hypertension 32 25391456
2010 Frequent inactivation of hSRBC in ovarian cancers by promoter CpG island hypermethylation. Acta obstetricia et gynecologica Scandinavica 24 20423276
2016 PRKCDBP (CAVIN3) and CRY2 associate with major depressive disorder. Journal of affective disorders 22 27721187
2019 ROR1-CAVIN3 interaction required for caveolae-dependent endocytosis and pro-survival signaling in lung adenocarcinoma. Oncogene 19 30894682
2012 Clinical relevance of loss of 11p15 in primary and metastatic breast cancer: association with loss of PRKCDBP expression in brain metastases. PloS one 17 23118876
2021 Cavin3 released from caveolae interacts with BRCA1 to regulate the cellular stress response. eLife 16 34142659
2014 Cavin-3 knockout mice show that cavin-3 is not essential for caveolae formation, for maintenance of body composition, or for glucose tolerance. PloS one 15 25036884
2023 The sodium new houttuyfonate suppresses NSCLC via activating pyroptosis through TCONS-14036/miR-1228-5p/PRKCDBP pathway. Cell proliferation 14 36696967
2012 CAVIN-3 regulates circadian period length and PER:CRY protein abundance and interactions. EMBO reports 14 23079727
2014 A Role for the Cavin-3/Matrix Metalloproteinase-9 Signaling Axis in the Regulation of PMA-Activated Human HT1080 Fibrosarcoma Cell Neoplastic Phenotype. Cancer growth and metastasis 10 25520561
2020 Matrine induces apoptosis and autophagy in human lung adenocarcinoma cells via upregulation of Cavin3 and suppression of PI3K/AKT pathway. Journal of B.U.ON. : official journal of the Balkan Union of Oncology 9 32862598
2017 Cavin-3 (PRKCDBP) deficiency reduces the density of caveolae in smooth muscle. Cell and tissue research 9 28285351
2021 Assessment of DAPK1 and CAVIN3 Gene Promoter Methylation in Breast Invasive Ductal Carcinoma and Metastasis. Cell journal 8 34455714
2020 Low expression of PRKCDBP promoted cisplatin resistance in lung adenocarcinoma by DNMT1 and TNF‑α. Oncology reports 7 32945503
2020 Cavin3 Suppresses Breast Cancer Metastasis via Inhibiting AKT Pathway. Frontiers in pharmacology 7 33101009
2014 Elevation of PRKCDBP, a novel transcriptional target of TNF-α, and its downregulation by infliximab in patients with ulcerative colitis. Digestive diseases and sciences 5 25052149
2020 Caveolae-Associated Protein 3 (Cavin-3) Influences Adipogenesis via TACE-Mediated Pref-1 Shedding. International journal of molecular sciences 4 32679831
2015 Polymorphisms in PRKCDBP, a Transcriptional Target of TNF-α, Are Associated With Inflammatory Bowel Disease in Korean. Intestinal research 4 26130999
2025 CAVIN3 deficiency promotes vascular normalization in ocular neovascular disease via ERK/JAG1 signaling pathway. JCI insight 3 40337864
2022 PRKCDBP Methylation is a Potential and Promising Candidate Biomarker for Non-small Cell Lung Cancer. Zhongguo fei ai za zhi = Chinese journal of lung cancer 3 35224960
2020 The Role of Cavin3 in the Progression of Lung Cancer and Its Mechanism. BioMed research international 3 32352004
2020 Inhibition of Cavin3 Degradation by the Human Parainfluenza Virus Type 2 V Protein Is Important for Efficient Viral Growth. Frontiers in microbiology 3 32425917
2025 The Role of Cavin3 in the Development of Malignant Tumors. Cancer management and research 1 40292077
2025 Cavin-3 promotes TNF expression via the MAPK signaling pathway in lung squamous cell carcinoma. Biochimica et biophysica acta. General subjects 0 41314262

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