Affinage

EOMES

Eomesodermin homolog · UniProt O95936

Length
686 aa
Mass
72.7 kDa
Annotated
2026-06-09
100 papers in source corpus 48 papers cited in narrative 48 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 5/5 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

EOMES (TBR2) is a T-box transcription factor that acts as a lineage-defining and stage-specific regulator across neural development, immune cell differentiation, and early embryonic germ-layer specification, operating largely by binding regulatory DNA and recruiting chromatin-modifying machinery (PMID:15634788, PMID:31792383, PMID:34521844). In the developing neocortex and hippocampus it marks the intermediate progenitor cell (IPC) state within a sequential Pax6→Tbr2→Tbr1 differentiation cascade and is both necessary and sufficient for IPC specification (PMID:15634788, PMID:18385329, PMID:23447624); it activates neuronal genes and represses stemness factors including Sox2, Pax6, and Insm1 by recruiting the H3K27me3 demethylase Jmjd3 (and partnering with NEUROG2) (PMID:22553033, PMID:27600842, PMID:30677129), and directly regulates downstream effectors of cortical circuit assembly such as Cxcl12-mediated interneuron migration and PCDH19-dependent clonal/synaptic organization (PMID:20713522, PMID:31477701). In the immune system EOMES is a lineage-defining factor for NK cells, driving early specification and maturation programs through direct enhancer binding, often in opposition to or alongside the paralog T-bet (PMID:22261438, PMID:34521844, PMID:38871999); in CD8+ T cells it controls memory, residency, and exhaustion programs, competing with T-bet at shared T-box sequences such as the Pdcd1 locus and directly driving prosurvival Bcl-2 and inhibitory/exhaustion genes including Havcr2, TIGIT, and CD226 loss (PMID:32182234, PMID:33979613, PMID:30619337, PMID:33053331), while in CD4+ T cells it induces IL-10, IFN-γ, and cytotoxic effector genes (PMID:30144030, PMID:36756120). In early embryogenesis EOMES cooperates with Brachyury to open mesoderm/endoderm-specific enhancers and drive ME fate while repressing default neuroectoderm, with EOMES specifically binding distal enhancers to specify anterior mesoderm and definitive endoderm in concert with SMAD2/3 and GATA6 (PMID:31792383, PMID:30629940, PMID:37633271, PMID:28174238), and instructing cardiogenic mesoderm via autocrine WNT3 signaling and a linc1405-tethered EOMES–WDR5–GCN5 complex (PMID:29382828, PMID:29754779). Across these contexts EOMES function converges on recruiting chromatin remodelers and modifiers—Jmjd3, BRG1, WDR5/GCN5—to reprogram enhancer accessibility at lineage-defining loci (PMID:30677129, PMID:31341159, PMID:29754779).

Mechanistic history

Synthesis pass · year-by-year structured walk · 25 steps
  1. 2005 High

    Established EOMES/Tbr2 as the molecular marker of the intermediate progenitor stage, resolving the order of transcription factors that drive cortical neuron production.

    Evidence Immunohistochemistry and marker co-expression in developing mouse neocortex defining a Pax6→Tbr2→Tbr1 cascade

    PMID:15634788

    Open questions at the time
    • Expression/lineage correlation does not establish direct transcriptional targets
    • Does not test necessity or sufficiency
  2. 2008 High

    Demonstrated that Tbr2 is necessary and sufficient for IPC identity, moving it from a marker to a causal specifier of cortical progenitor fate, and extended its role to adult hippocampal transit-amplifying cells.

    Evidence Forebrain-specific conditional knockout plus gain-of-function misexpression; adult hippocampal localization with neurogenic-stimulus manipulation

    PMID:18385329 PMID:18940588

    Open questions at the time
    • Direct transcriptional targets not yet defined
    • Mechanism of fate switch (activation vs repression) unresolved
  3. 2008 Medium

    Placed Tbr2 within an upstream signaling logic by showing Ngn2 directly activates it and Notch suppresses the cascade, explaining how the IPC program is gated during division.

    Evidence Single-cell live imaging in slice culture, gamma-secretase inhibition, direct target identification

    PMID:19059340

    Open questions at the time
    • Single lab
    • Asymmetric initiation mechanism not molecularly defined
  4. 2010 High

    Identified non-cell-autonomous EOMES outputs (Cxcl12) coordinating excitatory and inhibitory neuron balance, showing IPCs organize broader circuit assembly.

    Evidence Tbr2 conditional KO with Cxcl12 genetic rescue of interneuron migration

    PMID:20713522

    Open questions at the time
    • Whether Cxcl12 is a direct EOMES target not established here
    • Partial rescue implies additional effectors
  5. 2010 High

    Extended EOMES function beyond neural lineages to trophoblast stem cell self-renewal, the first evidence of context-specific transcriptional networks.

    Evidence ChIP-chip co-occupancy with TCFAP2C/SMARCA4 plus RNAi knockdown with transcriptome readout in mouse TS cells

    PMID:20176728

    Open questions at the time
    • ChIP-chip lower resolution than ChIP-seq
    • Direct vs indirect target distinction limited
  6. 2012 High

    Defined EOMES as a NK-cell lineage and maturation factor distinct from but partnered with T-bet, opening the immune arm of EOMES biology.

    Evidence Eomes/T-bet double conditional KO and stage-specific deletion in mature NK cells with flow cytometry staging

    PMID:22261438

    Open questions at the time
    • Direct NK target genes not yet mapped
    • Division of labor with T-bet not mechanistically resolved
  7. 2012 High

    Clarified the molecular basis of EOMES-driven neuronal lineage progression through Sox2 repression in the dentate gyrus and regional identity propagation in cortex.

    Evidence Conditional KO with Sox2 expression analysis (dentate gyrus); conditional inactivation with regional markers and axon tracing (cortex)

    PMID:22553033 PMID:23431145

    Open questions at the time
    • Whether Sox2 repression is direct not established here
    • Mechanism of regional gradient modulation unresolved
  8. 2013 High

    Connected EOMES expression to its chromatin/signaling control and downstream endoderm network, revealing an Activin–Jmjd3–Eomes positive feedback loop at a bivalent promoter.

    Evidence ChIP, chromatin looping, and Activin perturbation in differentiating mouse ES cells

    PMID:23584530

    Open questions at the time
    • Direct Eomes endoderm targets only partially mapped
    • Single differentiation system
  9. 2013 High

    Expanded EOMES neural roles to multiple neuron classes (olfactory bulb, retinal ipRGCs, hippocampal Cajal-Retzius migration) showing it controls excitatory-inhibitory balance and cell-type maintenance via Cxcr4 and other effectors.

    Evidence Conditional knockouts with anatomical, synaptic (EM), behavioral, and molecular (Cxcr4) readouts in mouse brain and retina

    PMID:22745484 PMID:23447624 PMID:25253855

    Open questions at the time
    • Direct targets in these neuron types largely undefined
    • Cell-autonomous vs non-autonomous contributions incompletely separated
  10. 2013 Medium

    Identified CREBBP as an EOMES partner and a repressive role at the IFNT promoter in ruminant trophoblast, an early protein-interaction and transcriptional-regulation finding.

    Evidence Co-IP and transient transfection reporter assays in bovine/ovine trophoblast cells

    PMID:23606646

    Open questions at the time
    • Ruminant system; relevance to mammalian EOMES generally unclear
    • Single lab co-IP without reciprocal validation
  11. 2014 High

    Positioned EOMES within the NK developmental transcriptional hierarchy (downstream of E4bp4, antagonized by T-bet), establishing that T-bet repression permits Eomes+ NK lineage emergence.

    Evidence ChIP showing E4bp4 binding the Eomes locus, genetic rescue of E4bp4-/- progenitors, Eomes-GFP reporter with ectopic T-bet expression

    PMID:24516120 PMID:24663216

    Open questions at the time
    • Mechanism of mutual T-bet/Eomes antagonism not yet at DNA-binding resolution
    • Microenvironmental cues controlling the switch incompletely defined
  12. 2016 High

    Defined an upstream Foxo3-Eomes axis and a CD4+ T cell program (non-classic Th1) in which Eomes promotes IFN-γ/cytotoxicity and represses ROR-γt/IL-17, extending EOMES to pathogenic effector differentiation.

    Evidence Foxo3 KO with lentiviral Eomes rescue in EAE; Eomes gain/loss in colitis model with cytokine profiling

    PMID:27742544 PMID:30144030

    Open questions at the time
    • Direct CD4 target genes not mapped here
    • Distinction from T-bet program incompletely defined
  13. 2017 High

    Provided genome-wide EOMES binding and a protein-interaction mechanism (NEUROG2, JMJD3) for the cortical IPC program, integrating genetic and epigenetic control.

    Evidence ChIP-seq, co-IP, and conditional KO expression profiling in mouse cortex

    PMID:27600842

    Open questions at the time
    • Functional contribution of each partner not dissected
    • Single lab
  14. 2018 High

    Established the embryonic enhancer logic and cardiogenic instructive capacity of EOMES, including a lincRNA-tethered EOMES-WDR5-GCN5 complex and autocrine WNT3 activation.

    Evidence Enhancer reporter/deletion with 3C and ATAC-seq; ChIP/RIP defining linc1405 complex; dose-dependent EOMES gain/loss in human ESCs with WNT manipulation

    PMID:28174238 PMID:29382828 PMID:29754779

    Open questions at the time
    • How EOMES recruits/uses each chromatin cofactor mechanistically incomplete
    • WNT shut-down switch regulation undefined
  15. 2019 High

    Resolved the core germ-layer decision: EOMES and Brachyury open ME enhancers and suppress default neuroectoderm, with EOMES uniquely specifying anterior mesoderm/definitive endoderm in concert with GATA6/SMAD2/3.

    Evidence Double KO in epiblast stem cells/embryos with ATAC-seq and ChIP-seq; genome-wide GATA6/EOMES/SMAD2/3 ChIP in human PSCs with patient-derived cells

    PMID:30629940 PMID:31792383

    Open questions at the time
    • Species differences (human vs mouse pancreatic ontogeny) not fully resolved
    • Cofactor switch determining anterior vs caudal fate not yet defined
  16. 2019 High

    Defined direct cortical effector genes (PCDH19) and consolidated the activate-PN/repress-stemness logic via Jmjd3 recruitment, and extended EOMES marking to spermatogonial stem cells.

    Evidence Conditional KO with MADM and PCDH19 rescue; ChIP-seq/co-IP review of Insm1/Pax6/Tbr1 regulation; lineage tracing and Plzf KO in SSCs

    PMID:30677129 PMID:31149899 PMID:31477701

    Open questions at the time
    • Review-sourced mechanistic detail (idx 24) attributed to primary studies
    • SSC transcriptional targets of EOMES undefined
  17. 2019 High

    Demonstrated EOMES recruits BRG1 chromatin remodeling at RUNX3-bound enhancers to establish innate memory CD8+ T cell programs, providing a direct epigenetic-reprogramming mechanism in T cells.

    Evidence ATAC-seq/H3K27ac ChIP-seq, co-IP of EOMES-BRG1 complex, BRG1 conditional KO in IL-4-driven innate memory model

    PMID:31341159

    Open questions at the time
    • Whether EOMES recruits BRG1 directly or via RUNX3 not fully resolved
    • Single context (innate memory)
  18. 2020 High

    Mapped EOMES direct transcriptional control of CD8+ T cell survival and inhibitory-receptor programs (Bcl-2, CD226, exhaustion genes) and identified upstream metabolic regulation via an AMPK-miR-107 axis.

    Evidence Conditional KO and ChIP identifying Bcl-2/CD226 targets with affinity-based and tumor models; AMPK/miR-107/Eomes pathway with PD-1 reporter and CAR-T model

    PMID:32182234 PMID:32221038 PMID:33053331

    Open questions at the time
    • Direction of inhibitory-receptor regulation context-dependent across studies
    • miR-107 axis (idx 28) single lab
  19. 2020 High

    Identified EOMES as a partner in lineage-specific trimeric complexes outside lymphoid cells (PU.1/MITF in osteoclasts) and proposed PTM-based regulation of its nuclear dynamics.

    Evidence Co-IP and sequential ChIP defining EOMES-PU.1-MITF complex with KD bone phenotype; antibody-based EOMES PTM detection in PD-1+ CD8 T cells

    PMID:30634169 PMID:32612611

    Open questions at the time
    • PTM mechanism (idx 32, Low confidence) lacks mutagenesis or enzyme assay validation
    • Osteoclast EOMES target genes not mapped
  20. 2021 High

    Resolved the EOMES/T-bet competition at the Pdcd1 T-box and defined subcellular-localization-dependent control of exhaustion, providing a mechanistic basis for differential PD-1 repression.

    Evidence Subcellular fractionation, Pdcd1 reporter competition assays, LCMV chronic infection and tumor models, human samples

    PMID:33979613

    Open questions at the time
    • What controls nuclear:cytoplasmic ratio of each factor incompletely defined
    • Single mechanistic context (exhaustion)
  21. 2021 High

    Established genome-wide that EOMES and T-bet bind largely overlapping NK targets with specificity conferred by differential cofactor recruitment, formalizing the sequential EOMES-early/T-bet-late maturation model.

    Evidence Endogenous EOMES and T-BET ChIP-seq, ATAC-seq, and conditional KO across NK stages

    PMID:34521844

    Open questions at the time
    • Identity of the differential cofactors not fully defined
    • Mechanism of motif-based discrimination from T-bet partly inferred
  22. 2021 Medium

    Expanded EOMES roles to CD8+ Treg localization, TRM lineage bifurcation, and direct binding of additional inhibitory loci (TIGIT, PD-1, CTLA-4, CD39) and a tumor-control requirement, broadening its immune regulatory map.

    Evidence Conditional KOs (TGF-βR/Eomes double KO, Hobit reporter), ChIP at inhibitory loci, siRNA rescue in patient-derived AML/HCC T cells, CLL tumor model

    PMID:30709927 PMID:32991667 PMID:33325636 PMID:33731848 PMID:34417257

    Open questions at the time
    • Direction of inhibitory-receptor regulation conflicts across tumor settings
    • Several findings single lab without reciprocal validation
  23. 2022 High

    Demonstrated tissue-compartment-specific EOMES functions in intestinal TRM, where it can both repress TRM formation and support maintenance depending on tissue, refining the context-dependent model.

    Evidence Conditional KO with transcriptomic and epigenetic profiling across small intestine and colon compartments

    PMID:36580919

    Open questions at the time
    • Molecular basis of tissue-specific outcome not defined
    • Direct targets driving the divergence unmapped
  24. 2023 High

    Provided the genome-wide mechanism of EOMES-Brachyury antagonism via differential distal enhancer binding, and showed in primary human NK cells that T-box factors suppress alternative ILC lineages.

    Evidence Endogenous Eomes/Brachyury ChIP-seq with single/double KO transcriptomes and motif analysis; CRISPR deletion in primary human NK cells with xenograft and scRNA-seq

    PMID:37279078 PMID:37633271

    Open questions at the time
    • How motif differences are read out by cofactors not fully resolved
    • Human-specific ILC-suppression targets incompletely mapped
  25. 2024 High

    Established EOMES as marking the bone-marrow NK-committed precursor distinct from PLZF+ ILC precursors, defining where the NK/ILC lineage split occurs.

    Evidence scRNA-seq and adoptive transfer into immunodeficient recipients with metastasis-prevention readout

    PMID:38871999

    Open questions at the time
    • Transcriptional targets locking in NK commitment not defined
    • How EOMES level sets the precursor identity unresolved

Open questions

Synthesis pass · forward-looking unresolved questions
  • How a single T-box factor achieves opposite outcomes (activation vs repression; lineage promotion vs antagonism) across neural, immune, and embryonic contexts remains unresolved at the level of cofactor combinatorics and post-translational control.
  • No unified model linking cofactor recruitment (Jmjd3, BRG1, WDR5/GCN5, PU.1/MITF, GATA6, SMAD2/3) to context-specific output
  • PTM-based regulation of EOMES nuclear activity not biochemically validated
  • Rules governing EOMES vs T-bet/Brachyury occupancy at shared loci incompletely defined

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0140110 transcription regulator activity 7 GO:0003677 DNA binding 5 GO:0140097 catalytic activity, acting on DNA 2
Localization
GO:0005634 nucleus 3
Pathway
R-HSA-1266738 Developmental Biology 5 R-HSA-168256 Immune System 5 R-HSA-4839726 Chromatin organization 4 R-HSA-74160 Gene expression (Transcription) 4
Complex memberships
EOMES-BRG1 chromatin complexEOMES-PU.1-MITF (osteoclast)EOMES-WDR5-GCN5 (linc1405-tethered)

Evidence

Reading pass · 48 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2005 Tbr2/EOMES expression defines the intermediate progenitor cell (IPC) stage in neocortical neurogenesis, marking the transition from Pax6+ radial glia to Tbr2+ IPCs to Tbr1+ postmitotic neurons, establishing a sequential transcription factor cascade Pax6 → Tbr2 → Tbr1 in the differentiation of radial glia → intermediate progenitor cell → postmitotic projection neuron. Immunohistochemistry, in situ hybridization, cell-type-specific marker co-expression in developing mouse neocortex The Journal of neuroscience High 15634788
2008 Conditional ablation of Tbr2/EOMES in the developing forebrain results in loss of intermediate progenitor cells (IPCs) and their neuronal progeny, reduced cortical surface expansion and thickness, and neuronal reduction across all cortical layers. Conversely, misexpression of Tbr2 in ventricular radial glial cells is sufficient to induce IPC identity, demonstrating Tbr2 is both necessary and sufficient for IPC specification. Conditional knockout mouse (forebrain-specific Tbr2 ablation), Tbr2::GFP transgenic cell fate tracing, gain-of-function misexpression Neuron High 18940588
2008 Tbr2/EOMES protein is specifically localized to intermediate-stage progenitor cells (IPCs) in the adult mouse hippocampus (subgranular zone), and Tbr2+ IPCs are highly responsive to neurogenic stimuli (voluntary wheel running more than doubles their number), establishing Tbr2 as a marker of transit-amplifying cells in adult hippocampal neurogenesis operating within the same Pax6 → Ngn2 → Tbr2 → NeuroD → Tbr1 transcription factor cascade as embryonic cortex. Immunostaining, Tbr2-GFP transgene expression, BrdU/antimitotic drug manipulation, multi-TF cascade analysis in adult mouse brain The Journal of neuroscience High 18385329
2008 Ngn2 directly activates Tbr2 transcription in nascent neocortical daughter cells; Notch signaling in periventricular daughter cells suppresses this Ngn2-Tbr2 cascade. Tbr2 expression is initiated asymmetrically in one of the two daughter cells following radial glial division, biased toward the apical cell. Single-cell DiI labeling and time-lapse tracking in slice cultures, immunostaining, gamma-secretase inhibition, identification of Tbr2 as direct Ngn2 target Molecular and cellular neurosciences Medium 19059340
2010 Tbr2-positive cortical intermediate progenitors control the amount and migratory route of subpallial GABAergic interneurons into the subventricular zone through a non-cell-autonomous mechanism mediated by Cxcl12 chemokine signaling. Forced Cxcl12 expression in Tbr2 mutants partially rescues SVZ interneuron migration, placing Tbr2 upstream of Cxcl12 in coordinating excitatory and inhibitory neuron balance. Tbr2 conditional knockout, interneuron migration analysis, Cxcl12 rescue experiment in mutant cortex Genes & development High 20713522
2010 EOMES co-occupies a significant number of genomic loci with TCFAP2C and SMARCA4 in trophoblast stem (TS) cells as determined by ChIP-chip. RNAi depletion of Eomes leads to loss of normal TS cell colony morphology and downregulation of TS cell-specific genes, establishing EOMES as a component of the transcriptional network required for TS cell self-renewal. ChIP-chip (chromatin immunoprecipitation with microarray), RNAi knockdown, transcriptome analysis in mouse trophoblast stem cells Genome research High 20176728
2012 Mice lacking both Eomes and T-bet fail to develop NK cells. Eomes is specifically required for the maturation step characterized by loss of constitutive TRAIL expression and induction of Ly49 receptor diversity and CD49b (DX5+), while T-bet governs developmental stability of immature TRAIL+ NK cells. Deletion of Eomes from mature NK cells causes reversion to phenotypic immaturity. Eomes/T-bet double conditional knockout mice, NK cell developmental stage analysis by flow cytometry, genetic deletion of Eomes in mature NK cells Immunity High 22261438
2012 Tbr2/EOMES is critically required for neurogenesis in the dentate gyrus: in the absence of Tbr2, intermediate neuronal progenitors (INPs) are depleted despite augmented neural stem cell proliferation, neurogenesis is halted due to failed neuronal differentiation, and Tbr2 likely promotes lineage progression from NSC to INP in part by repression of Sox2. Conditional Tbr2 knockout in mouse dentate gyrus, flow cytometry, immunostaining, Sox2 expression analysis The Journal of neuroscience High 22553033
2013 Tbr2/EOMES is required for proper migration of Cajal-Retzius cells to the dentate gyrus (DG), formation of the hippocampal fissure, and establishment of the transhilar radial glial scaffold. Loss of Tbr2 results in decreased Cxcr4 expression in migrating cells, leading to premature granule neurogenesis and increased cell death, and depletes the DG stem cell population before proper SGZ establishment. Conditional Tbr2 knockout, cell migration analysis, Cxcr4 expression profiling, BrdU labeling in developing mouse dentate gyrus The Journal of neuroscience High 23447624
2013 The histone demethylase Jmjd3 sequentially associates with Tbx3 (at the Eomes enhancer) and then with Eomes itself (at its own bivalent promoter) to drive Eomes expression during endodermal differentiation of embryonic stem cells. Eomes then activates a transcriptional network of core regulators of definitive endoderm differentiation, and Activin signaling promotes Jmjd3/Eomes binding at the Eomes promoter in a positive feedback loop. ChIP, chromatin conformation/looping analysis, Activin signaling perturbation, gene expression analysis in differentiating mouse ES cells The EMBO journal High 23584530
2013 Eomes inactivation in the cortical subventricular zone causes rostrocaudal shifts in SVZ and CP gene expression gradients (the 'intermediate map'), with loss of corticospinal axons and gain of corticotectal projections, establishing Eomes as a modulator of regional identity propagation from VZ progenitors to cortical plate neurons via the Pax6 → Eomes → Tbr1 cascade. Conditional Eomes inactivation, in situ hybridization for regional markers, axonal tract tracing in mouse neocortex Proceedings of the National Academy of Sciences of the United States of America High 23431145
2013 Tbr2 deficiency in olfactory bulb mitral and tufted cells causes cell-autonomous changes including compensatory upregulation of Tbr1 and VGluT subtype shift (VGluT1 to VGluT2), dendritic morphology and projection abnormalities, non-cell-autonomous loss of inhibitory interneuron subtypes, reduction in dendrodendritic reciprocal synapses, and hyperactivation of mitral/tufted cells by odorants—establishing Tbr2 as required for excitatory-inhibitory balance in the olfactory bulb. Conditional Tbr2 knockout in mitral/tufted cells, immunostaining, odorant stimulation/c-Fos analysis, electron microscopy for synapse counting The Journal of neuroscience High 22745484
2014 E4bp4/Nfil3 directly binds to the regulatory regions of both Eomes and Id2 and promotes their transcription, acting upstream of Eomes in NK lineage commitment. Eomes and Id2 can rescue NK production from E4bp4-/- progenitors, placing Eomes downstream of E4bp4 in the NK development transcriptional hierarchy. ChIP, genetic rescue of E4bp4-/- NK progenitors with Eomes/Id2 overexpression, E4bp4 knockout mouse The Journal of experimental medicine High 24663216
2014 In the liver, T-bet expression in progenitors represses Eomes expression and the development of Eomes+ NK cells. Conversely, the bone marrow microenvironment restricts T-bet expression in developing NK cells, permitting Eomes-dependent NK maturation. Ectopic expression of T-bet forces development of Eomes- NK cells, demonstrating that T-bet repression is essential for Eomes+ NK cell development. Eomes- and Eomes+ NK cells constitute distinct lineages with complementary functions. Eomes-GFP reporter mouse generation, ectopic T-bet expression, gene expression profiling, Eomes conditional knockout analysis The Journal of experimental medicine High 24516120
2014 Tbr2/EOMES is essential for the formation and maintenance of melanopsin (Opn4)-expressing intrinsically photosensitive retinal ganglion cells (ipRGCs): Opn4 expression is exclusive to Tbr2+ RGCs, genetic ablation of Tbr2 before RGC specification eliminates all ipRGCs, and deletion of Tbr2 in established ipRGCs eliminates most of them and causes dendritic morphology defects. Conditional Tbr2 knockout (pre- and post-specification), single-cell dye tracing, behavioral analysis, immunostaining in mouse retina The Journal of neuroscience High 25253855
2016 Eomes controls non-classic Th1 (Th17-derived Th1) cell development by promoting IFN-γ secretion while inhibiting ROR-γt expression and IL-17 production. In a mouse model of T cell-dependent colitis, Eomes drives non-classic Th1 pathogenic potential in vivo; Eomes also promotes acquisition of a cytotoxic signature and IFN-γ+GM-CSF+ cell development. Eomes overexpression/conditional knockout in colitis mouse model, flow cytometry, cytokine analysis, ROR-γt expression analysis European journal of immunology Medium 30144030
2016 Foxo3 directly targets and activates the Eomes gene in CD4+ T cells. Lentiviral overexpression of Eomes in Foxo3-deficient CD4+ T cells restores IFN-γ and GM-CSF production, establishing a Foxo3-Eomes axis required for pathogenic Th1 cell differentiation and neuroinflammation. Foxo3 knockout mice, EAE model, lentiviral Eomes overexpression in Foxo3-deficient T cells, direct Eomes target gene identification Immunity High 27742544
2017 TBR2 directly regulates a molecular network in intermediate neuronal progenitors (INPs) through protein-protein interactions with NEUROG2 and JMJD3, revealing both genetic and epigenetic mechanisms. Genome-wide ChIP-seq identified TBR2 direct target genes, and co-IP confirmed NEUROG2 and JMJD3 as TBR2 binding partners controlling INP identity, morphology, proliferation, and migration. ChIP-seq (TBR2 genome-wide binding), global gene expression profiling, co-immunoprecipitation, conditional knockout analysis in mouse cortex Cerebral cortex High 27600842
2017 A visceral/primitive streak enhancer element (VPE) in the Eomes locus is required for optimal Eomes expression in vivo; deletion of VPE causes variable defects in anterior-posterior axis orientation and definitive endoderm formation. Chromosome conformation capture shows VPE-promoter interactions in ES cells prior to gene activation; activation during DE differentiation is associated with Smad2/3 binding and increased chromatin accessibility at VPE, occurring without large-scale 3D chromatin reorganization. Transgenic enhancer reporter assays, targeted enhancer deletion, chromosome conformation capture (3C/4C), ATAC-seq, Smad2/3 ChIP in mouse embryos and ES cells Development High 28174238
2018 A lincRNA (linc1405) physically mediates a complex consisting of Eomes, TrxG subunit WDR5, and histone acetyltransferase GCN5 that binds the enhancer region of the Mesp1 gene to activate its expression during cardiac mesoderm specification of embryonic stem cells. Linc1405 depletion impairs heart development and function in vivo. Chromatin immunoprecipitation (ChIP), RNA immunoprecipitation (RIP), linc1405 knockdown in ESCs and in vivo zebrafish/mouse models Cell stem cell High 29754779
2018 Dose-dependent induction of EOMES alone can fully replace a cocktail of signaling molecules for cardiogenic mesoderm specification from human embryonic stem cells. Mechanistically, EOMES-driven cardiomyocyte programming involves autocrine activation of canonical WNT signaling via WNT3 ligand, which must subsequently be shut down for cardiomyocyte maturation. EOMES gain- and loss-of-function in human ESCs, directed differentiation assays, WNT pathway inhibition/activation experiments Nature communications High 29382828
2019 Eomes and Brachyury are required for mesoderm and definitive endoderm (ME) gene programs; cells deficient in these T-box factors retain pluripotency and default to neuroectoderm lineages even in the presence of TGF-β/Nodal and Wnt ME-inducing signals. Eomes and Brachyury bind ME enhancers and are required for their chromatin accessibility, whereas NE enhancers are pre-accessible. Eomes uniquely specifies anterior mesoderm and definitive endoderm, while Brachyury specifies caudal mesoderm. Conditional double KO of Eomes and Brachyury in mouse epiblast stem cells and embryos, ATAC-seq (chromatin accessibility), ChIP-seq, transcriptome analysis Nature cell biology High 31792383
2019 TBR2 directly regulates Protocadherin 19 (PCDH19) expression, and PCDH19 mediates TBR2's role in coordinating clonal organization and preferential synapse development of excitatory neurons. Simultaneous PCDH19 expression rescues both neurogenesis and neuronal organization defects caused by TBR2 removal, placing PCDH19 as a direct downstream effector of TBR2 in cortical microcircuit assembly. Conditional TBR2 knockout, mosaic analysis with double markers (MADM), PCDH19 rescue experiments, electrophysiology in mouse cortex Nature communications High 31477701
2019 GATA6 binds and cooperates with EOMES/SMAD2/3 to regulate expression of cardinal definitive endoderm genes in human pluripotent stem cells, as established by genome-wide ChIP studies. GATA6-null hPSCs fail to enter the DE lineage, an effect recapitulated in patient-derived cells, demonstrating that the GATA6-EOMES/SMAD2/3 axis is specifically required for human (but not mouse) pancreatic ontogeny. Genome-wide ChIP (GATA6, EOMES, SMAD2/3), GATA6 gene-edited and patient-derived hPSCs, directed differentiation toward DE and β-cells Stem cell reports High 30629940
2019 Tbr2/EOMES directly represses Insm1 (an IP-genic transcription factor gene) and Pax6 (a key activator of Tbr2 transcription), and activates PN-specific genes such as Tbr1 by recruiting Jmjd3, a histone H3K27me3 demethylase that removes repressive epigenetic marks placed by polycomb repressive complex 2. ChIP-seq (Tbr2 direct targets), Jmjd3 co-IP/recruitment assay, Tbr2 conditional KO transcriptome analysis, H3K27me3 histone mark profiling Journal of anatomy Medium 30677129
2019 EOMES is expressed in a rare subpopulation of spermatogonial stem cells (SSCs). Lineage tracing and busulfan challenge demonstrate these EOMES+ cells are SSCs contributing to steady-state spermatogenesis and regeneration following injury. PLZF regulates the proliferative activity of EOMES+ SSCs: EOMES+ SSCs have lower proliferation index in wild-type than in Plzf-/- mice, and are lost through proliferative exhaustion in Plzf-/- mice. Lineage tracing (Eomes-CreERT2), busulfan chemical injury model, RNAseq, single-cell RNAseq, Plzf knockout mice eLife High 31149899
2019 EOMES interacts with RUNX3-bound enhancers and recruits BRG1 (a chromatin remodeling factor) to drive epigenetic reprogramming establishing the innate memory CD8+ T cell program. EOMES is found within chromatin-associated complexes containing BRG1, and the in vivo acquisition of the EOMES-dependent program is BRG1-dependent. Epigenomic profiling (ATAC-seq, H3K27ac ChIP-seq), co-immunoprecipitation identifying EOMES-BRG1 complex, BRG1 conditional KO, IL-4-driven innate memory model Nature communications High 31341159
2020 Accumulation of CD226neg CD8+ T cells in tumors is driven by an antigen-specific mechanism involving EOMES. EOMES directly drives CD226 loss in CD8+ T cells, and anti-CD137 agonists stimulate Eomes-dependent CD226 downregulation that limits anti-tumor efficacy. CD226 loss is associated with reduced LFA-1 activation and altered TCR signaling. Flow cytometry in human/mouse tumors, Eomes conditional KO and overexpression, ChIP/ChIP-seq, anti-PD-1 and anti-CD137 treatment models Immunity High 33053331
2020 AMPK activation by metformin decreases microRNA-107 expression, which enhances Eomesodermin expression, which in turn suppresses PDCD1 (PD-1) transcription in CD8+ T cells, promoting memory T cell formation via an AMPK-miR-107-Eomes-PD-1 pathway. In vitro metformin treatment, AMPK activation assay, miR-107 expression analysis, Eomes overexpression/knockdown, PD-1 promoter reporter assay, CAR-T tumor model Journal of immunology Medium 32221038
2020 EOMES partners with PU.1 and MITF in a trimeric complex at genomic loci critical for osteoclast differentiation. Sequential ChIP confirmed co-occupancy. EOMES knockdown in myeloid precursors leads to osteopetrosis with decreased osteoclast differentiation and function in vitro and in vivo. Co-immunoprecipitation, sequential ChIP (re-ChIP), EOMES knockdown in myeloid precursors, in vivo mouse bone phenotype analysis iScience High 30634169
2020 Eomes directly drives transcription of the prosurvival protein Bcl-2 in CD8+ T cells of low activation signal strength; this is induced at low signal intensity while T-bet is induced at higher signal intensity and suppresses Bcl-2. The Eomes/Bcl-2 axis preserves low-affinity CD8+ T cell clones in memory, maintaining clonal diversity. Eomes conditional KO, ChIP identifying Bcl-2 as direct Eomes target, affinity-based T cell activation models, clonal diversity analysis PLoS biology High 32182234
2020 Stage-specific deletion of Eomes in mature NK cells (using tamoxifen-inducible Ncr1-targeted Cre) results in rapid loss of NK cells with particularly profound depletion of penultimately mature stage III NK cells, through increased apoptosis and impaired maturation from stage II precursors. Induced Eomes deletion also decreases NK cell cytotoxicity and abrogates in vivo rejection of MHC-class-I-deficient cells. Tamoxifen-inducible Ncr1-Cre × Eomes-flox conditional deletion, NK cell staging by flow cytometry, in vivo NK cell rejection assay, apoptosis assays Cell reports High 32492428
2020 Nuclear LSD1 phosphorylated at serine 111 (nLSD1p) regulates EOMES nuclear dynamics via demethylation/acetylation switching of critical EOMES residues. EOMES co-exists with nLSD1p in PD-1+CD8+ T cells, and EOMES demethylation/acetylation is reciprocally expressed in immunotherapy-resistant versus responder patients. Novel antibodies detecting EOMES PTMs, co-expression analysis, nLSD1 inhibitor treatment, flow cytometry in patient samples and 4T1 mouse model Frontiers in immunology Low 32612611
2021 The subcellular localization of T-bet and Eomes dictates their regulatory activity in exhausted T cells (TEXs): TEXs have a higher nuclear Eomes:T-bet ratio than memory T cells. T-bet and Eomes compete for the same DNA sequences including the Pdcd1 T-box; high nuclear T-bet strongly represses Pdcd1 transcription whereas low nuclear T-bet in TEX leads to dominant Eomes acting as a weaker repressor of Pdcd1. PD-1 blockade increases nuclear T-bet, restoring stronger Pdcd1 repression. Subcellular fractionation, nuclear/cytoplasmic T-bet and Eomes quantification, Pdcd1 reporter assays with T-bet/Eomes competition, LCMV chronic infection model, preclinical cancer models, human tumor samples Cell reports High 33979613
2021 EOMES sequentially regulates NK cell development by driving early lineage specification and hallmark receptor/function induction in immature NK cells, while T-BET dominates in mature NK cells by inducing IL-12 responsiveness and repressing the cell cycle. ChIP of endogenous EOMES and T-BET shows strong overlap in DNA-binding targets with specificity mediated by differential co-factor recruitment and epigenetic changes during differentiation. Gene-modified mice enabling ChIP of endogenous EOMES and T-BET, ChIP-seq, ATAC-seq, Eomes/T-bet conditional KO mice, NK cell staging Nature communications High 34521844
2021 Eomes controls the follicular location of CD8+ regulatory T cells (CD8+ Tregs), while TGF-β receptor signaling maintains their regulatory identity. Simultaneous disruption of TGF-β receptor and Eomes in T cells causes lethal autoimmunity due to specific loss of CD8+ (but not CD4+) Tregs. Conditional double KO of TGF-β receptor and Eomes in T cells, autoimmunity phenotype analysis, CD8+ vs CD4+ Treg quantification, tissue localization The Journal of experimental medicine High 32991667
2021 EOMES directly bound the promoter of T-cell immunoglobulin and ITIM domain (TIGIT) and positively regulated TIGIT expression on patient-derived T cells. siRNA-mediated depletion of Eomes reversed functional defects (cytokine production and killing capacity) in Eomes+T-betlow CD8+ T cells from AML patients. ChIP (EOMES binding to TIGIT promoter), siRNA knockdown, flow cytometry functional assays in patient-derived AML T cells Cancer research Medium 30709927
2021 Eomes is identified as a key factor instructing early bifurcation of circulating and tissue-resident (TRM) CD8+ T cell lineages: Hobit+ TRM precursors form during the effector phase, and Eomes regulates the early commitment to residency versus circulation. Hobit reporter/deleter mice, Eomes conditional KO, adoptive transfer of Hobit+ effector cells, transcriptional profiling of TRM precursors Science immunology Medium 34417257
2021 EOMES-deficient CD8+ T cells show impaired expansion and decreased CLL tumor control in vivo. EOMES is therefore essential for CD8+ T cell expansion/maintenance required for adaptive immune control of CLL. Eomes conditional KO in CD8+ T cells, Eµ-TCL1 CLL mouse model, tumor control assay, flow cytometry Leukemia Medium 33731848
2021 Eomes directly bound the transcriptional regulatory regions of the key inhibitory receptors PD-1, CTLA-4, and CD39 in CD8+ T cells from hepatocellular carcinoma tumors; lower EOMES levels in tumor-infiltrating CD8+ T cells contributed to overexpression of these inhibitory factors. ChIP (EOMES binding to PD-1, CTLA-4, CD39 regulatory regions), flow cytometry in patient tumor-infiltrating lymphocytes Journal of cellular and molecular medicine Medium 33325636
2021 Eomes directly controls expression of T cell exhaustion gene Havcr2 (Tim-3) through high-level Eomes binding to regulatory genomic loci, and Eomes may compete with T-bet at shared regulatory genomic loci to antagonize T-bet functions. Integrated RNAseq and ChIPseq in Eomes-overexpressing T cells revealed direct Eomes transcriptional control of exhaustion genes. ChIP-seq (Eomes binding), RNAseq in Eomes-overexpressing T cells, Eomes heterozygous deletion in tumor model, Havcr2 direct target validation Frontiers in immunology Medium 30619337
2019 Extrapituitary prolactin (PRL) produced by B cells and MHC class II+ myeloid cells in late EAE lesions induces Eomes+ CD4+ T (Th) cells from naïve T cells ex vivo in an MHC class II-dependent manner. Blocking PRL production (bromocriptine or Zbtb20-specific siRNA) significantly reduced Eomes+ Th cells in the CNS and ameliorated late EAE clinical signs. EAE mouse model, PRL blocking (bromocriptine, Zbtb20 siRNA), ex vivo PRL stimulation, anti-MHC class II antibody blockade Proceedings of the National Academy of Sciences of the United States of America Medium 31570595
2022 Eomes exhibits context-specific roles in intestinal tissue-resident memory CD8+ T cells (TRM): while Eomes represses TRM formation in some tissues, it supports maintenance of established TRM cells in the small intestine but not in the colon, revealing tissue-compartment-specific regulatory functions. Eomes conditional KO, TRM isolation from lamina propria and epithelial compartments of small intestine and colon, transcriptomic and epigenetic profiling Immunity High 36580919
2023 On a genome-wide scale, Eomes and Brachyury binding overlaps at target gene promoters but shows specificity for distal enhancer regions conferred by differences in Tbx DNA-binding motifs. Eomes antagonizes Brachyury gene regulatory functions in coexpressing cells during early gastrulation via these differential enhancer binding preferences, ensuring proper sequence of anterior mesoderm/DE followed by posterior mesoderm specification. ChIP-seq (endogenous Eomes and Brachyury), Eomes/Brachyury single and double KO in mouse embryos and stem cells, transcriptome analysis, enhancer motif analysis Developmental cell High 37633271
2023 Deletion of T-BET and EOMES in primary human NK cells using CRISPR/Cas9 compromises in vivo antitumor response, NK cell proliferation and persistence, and cytokine responses. T-BET and EOMES deletion causes CD56bright NK cells to acquire an innate lymphoid cell precursor-like (ILCP-like) profile with increased RORC and AHR (ILC3-associated TFs), demonstrating that T-box TFs suppress alternative ILC lineages in mature NK cells. CRISPR/Cas9 deletion in primary human NK cells, in vivo xenograft tumor models, single-cell RNA-seq, TF expression analysis The Journal of clinical investigation High 37279078
2023 Eomes expression in murine CD4+ T cells is sufficient to induce IL-10 expression and, together with T-bet, promotes a cytotoxic effector profile (Prf1, Gzmb, Gzmk, Nkg7, Ccl5) while repressing alternative cell fates (Th2, Th17). The Eomes-driven transcriptional program in CD4+ T cells only partially overlaps with that of T-bet. Novel genetic approach to enforce Eomes expression in CD4+ T cells, transcriptomic profiling (RNA-seq), cytokine/effector molecule measurement by flow cytometry Frontiers in immunology Medium 36756120
2024 EomeshiNKneg cells in the bone marrow (lineage-negative, NK receptor-negative, Eomes-expressing) are the bone marrow precursors to classical NK cells. Transfer of EomeshiNKneg cells into Rag2-/-Il2rg-/- recipients generates functional NK cells capable of preventing metastatic disease, while PLZF+ ILC precursors generate ILC1s, ILC2s, and ILC3s lacking cytotoxic potential—establishing that NK and ILC1 lineages diverge early in development and that Eomes marks the NK-committed precursor. Single-cell RNA-seq, flow cytometry, adoptive transfer into immunodeficient recipients, metastasis prevention assay Nature immunology High 38871999
2013 EOMES interacts with CREBBP (CBP) as shown by immunoprecipitation, and EOMES down-regulates interferon tau (IFNT) gene transcription in bovine/ovine trophoblast cells. Transient transfection reporter assays showed EOMES suppresses IFNT promoter transactivation. Uterine flushing from pregnant animals increases EOMES expression in bovine trophoblast CT-1 cells specifically in co-culture with uterine epithelial cells. Co-immunoprecipitation (EOMES-CREBBP), transient transfection reporter assay, real-time PCR, CT-1/uterine epithelial cell co-culture Molecular reproduction and development Medium 23606646

Source papers

Stage 0 corpus · 100 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2005 Pax6, Tbr2, and Tbr1 are expressed sequentially by radial glia, intermediate progenitor cells, and postmitotic neurons in developing neocortex. The Journal of neuroscience : the official journal of the Society for Neuroscience 1069 15634788
2012 The transcription factors T-bet and Eomes control key checkpoints of natural killer cell maturation. Immunity 624 22261438
2014 T-bet and Eomes instruct the development of two distinct natural killer cell lineages in the liver and in the bone marrow. The Journal of experimental medicine 471 24516120
2008 Tbr2 directs conversion of radial glia into basal precursors and guides neuronal amplification by indirect neurogenesis in the developing neocortex. Neuron 308 18940588
2008 Intermediate progenitors in adult hippocampal neurogenesis: Tbr2 expression and coordinate regulation of neuronal output. The Journal of neuroscience : the official journal of the Society for Neuroscience 257 18385329
2014 T-bet and Eomes are differentially linked to the exhausted phenotype of CD8+ T cells in HIV infection. PLoS pathogens 249 25032686
2016 CXCR6 marks a novel subset of T-bet(lo)Eomes(hi) natural killer cells residing in human liver. Scientific reports 211 27210614
2018 High Levels of Eomes Promote Exhaustion of Anti-tumor CD8+ T Cells. Frontiers in immunology 174 30619337
2014 Characterization of T-bet and eomes in peripheral human immune cells. Frontiers in immunology 167 24860576
2014 The transcription factor E4bp4/Nfil3 controls commitment to the NK lineage and directly regulates Eomes and Id2 expression. The Journal of experimental medicine 159 24663216
2007 Homozygous silencing of T-box transcription factor EOMES leads to microcephaly with polymicrogyria and corpus callosum agenesis. Nature genetics 155 17353897
2018 T-bet and Eomes govern differentiation and function of mouse and human NK cells and ILC1. European journal of immunology 143 29424438
2014 Cortical and Clonal Contribution of Tbr2 Expressing Progenitors in the Developing Mouse Brain. Cerebral cortex (New York, N.Y. : 1991) 135 24927931
1999 Expression pattern of the Tbr2 (Eomesodermin) gene during mouse and chick brain development. Mechanisms of development 124 10473127
2020 Metformin Enhances the Antitumor Activity of CD8+ T Lymphocytes via the AMPK-miR-107-Eomes-PD-1 Pathway. Journal of immunology (Baltimore, Md. : 1950) 123 32221038
2016 Tissue-resident Eomes(hi) T-bet(lo) CD56(bright) NK cells with reduced proinflammatory potential are enriched in the adult human liver. European journal of immunology 123 27485474
2020 Eomes-Dependent Loss of the Co-activating Receptor CD226 Restrains CD8+ T Cell Anti-tumor Functions and Limits the Efficacy of Cancer Immunotherapy. Immunity 122 33053331
2012 Tbr2 is essential for hippocampal lineage progression from neural stem cells to intermediate progenitors and neurons. The Journal of neuroscience : the official journal of the Society for Neuroscience 119 22553033
2004 Expression of the genes Emx1, Tbr1, and Eomes (Tbr2) in the telencephalon of Xenopus laevis confirms the existence of a ventral pallial division in all tetrapods. The Journal of comparative neurology 118 15174073
2010 Examination of transcriptional networks reveals an important role for TCFAP2C, SMARCA4, and EOMES in trophoblast stem cell maintenance. Genome research 111 20176728
2013 Differential localization of T-bet and Eomes in CD8 T cell memory populations. Journal of immunology (Baltimore, Md. : 1950) 109 23455505
2016 Unique Eomes(+) NK Cell Subsets Are Present in Uterus and Decidua During Early Pregnancy. Frontiers in immunology 103 27004067
2019 Intermediate progenitors and Tbr2 in cortical development. Journal of anatomy 100 30677129
2021 Role of nuclear localization in the regulation and function of T-bet and Eomes in exhausted CD8 T cells. Cell reports 97 33979613
2012 Diagnosis of bladder cancer recurrence based on urinary levels of EOMES, HOXA9, POU4F2, TWIST1, VIM, and ZNF154 hypermethylation. PloS one 97 23056278
2019 Eomes and Brachyury control pluripotency exit and germ-layer segregation by changing the chromatin state. Nature cell biology 86 31792383
2010 Tbr2-positive intermediate (basal) neuronal progenitors safeguard cerebral cortex expansion by controlling amplification of pallial glutamatergic neurons and attraction of subpallial GABAergic interneurons. Genes & development 86 20713522
2021 Sequential actions of EOMES and T-BET promote stepwise maturation of natural killer cells. Nature communications 83 34521844
2018 A Linc1405/Eomes Complex Promotes Cardiac Mesoderm Specification and Cardiogenesis. Cell stem cell 83 29754779
2008 Periventricular notch activation and asymmetric Ngn2 and Tbr2 expression in pair-generated neocortical daughter cells. Molecular and cellular neurosciences 83 19059340
2013 The histone demethylase Jmjd3 sequentially associates with the transcription factors Tbx3 and Eomes to drive endoderm differentiation. The EMBO journal 78 23584530
2021 Hobit identifies tissue-resident memory T cell precursors that are regulated by Eomes. Science immunology 77 34417257
2013 The protomap is propagated to cortical plate neurons through an Eomes-dependent intermediate map. Proceedings of the National Academy of Sciences of the United States of America 77 23431145
2020 Eomes identifies thymic precursors of self-specific memory-phenotype CD8+ T cells. Nature immunology 71 32284593
2021 Involvement of cytotoxic Eomes-expressing CD4+ T cells in secondary progressive multiple sclerosis. Proceedings of the National Academy of Sciences of the United States of America 65 33836594
2019 TBR2 coordinates neurogenesis expansion and precise microcircuit organization via Protocadherin 19 in the mammalian cortex. Nature communications 65 31477701
2016 Prenatal β-catenin/Brn2/Tbr2 transcriptional cascade regulates adult social and stereotypic behaviors. Molecular psychiatry 65 26830142
2013 Tbr2 expression in Cajal-Retzius cells and intermediate neuronal progenitors is required for morphogenesis of the dentate gyrus. The Journal of neuroscience : the official journal of the Society for Neuroscience 64 23447624
2022 Small intestine and colon tissue-resident memory CD8+ T cells exhibit molecular heterogeneity and differential dependence on Eomes. Immunity 63 36580919
2019 Identification of EOMES-expressing spermatogonial stem cells and their regulation by PLZF. eLife 61 31149899
2009 Generation and analysis of a mouse line harboring GFP in the Eomes/Tbr2 locus. Genesis (New York, N.Y. : 2000) 61 19830823
2008 Progressive loss of PAX6, TBR2, NEUROD and TBR1 mRNA gradients correlates with translocation of EMX2 to the cortical plate during human cortical development. The European journal of neuroscience 61 18973570
2005 Differential transcription of Eomes and T-bet during maturation of mouse uterine natural killer cells. Journal of leukocyte biology 61 16204645
2007 Eomes::GFP-a tool for live imaging cells of the trophoblast, primitive streak, and telencephalon in the mouse embryo. Genesis (New York, N.Y. : 2000) 60 17417802
2018 Eomes controls the development of Th17-derived (non-classic) Th1 cells during chronic inflammation. European journal of immunology 59 30144030
2021 TGF-β and Eomes control the homeostasis of CD8+ regulatory T cells. The Journal of experimental medicine 58 32991667
2020 Stage-Specific Requirement for Eomes in Mature NK Cell Homeostasis and Cytotoxicity. Cell reports 58 32492428
2013 T-Bet and Eomes Regulate the Balance between the Effector/Central Memory T Cells versus Memory Stem Like T Cells. PloS one 58 23826287
2016 Foxo3 Transcription Factor Drives Pathogenic T Helper 1 Differentiation by Inducing the Expression of Eomes. Immunity 57 27742544
2021 Clonally expanded EOMES+ Tr1-like cells in primary and metastatic tumors are associated with disease progression. Nature immunology 55 34017124
2014 T-box transcription regulator Tbr2 is essential for the formation and maintenance of Opn4/melanopsin-expressing intrinsically photosensitive retinal ganglion cells. The Journal of neuroscience : the official journal of the Society for Neuroscience 54 25253855
2020 Targeting Nuclear LSD1 to Reprogram Cancer Cells and Reinvigorate Exhausted T Cells via a Novel LSD1-EOMES Switch. Frontiers in immunology 52 32612611
1999 A novel mammalian T-box-containing gene, Tbr2, expressed in mouse developing brain. Brain research. Developmental brain research 52 10407135
2023 T-BET and EOMES sustain mature human NK cell identity and antitumor function. The Journal of clinical investigation 49 37279078
2019 Eomes+T-betlow CD8+ T Cells Are Functionally Impaired and Are Associated with Poor Clinical Outcome in Patients with Acute Myeloid Leukemia. Cancer research 49 30709927
2014 Integrated analyses of DNA methylation and hydroxymethylation reveal tumor suppressive roles of ECM1, ATF5, and EOMES in human hepatocellular carcinoma. Genome biology 49 25517360
2014 The autoimmune disease-associated transcription factors EOMES and TBX21 are dysregulated in multiple sclerosis and define a molecular subtype of disease. Clinical immunology (Orlando, Fla.) 48 24495857
2019 Tissue-resident Eomes+ NK cells are the major innate lymphoid cell population in human infant intestine. Nature communications 47 30816112
1999 Identification, mapping, and phylogenomic analysis of four new human members of the T-box gene family: EOMES, TBX6, TBX18, and TBX19. Genomics 47 9888994
2019 EOMES interacts with RUNX3 and BRG1 to promote innate memory cell formation through epigenetic reprogramming. Nature communications 46 31341159
2021 EOMES is essential for antitumor activity of CD8+ T cells in chronic lymphocytic leukemia. Leukemia 44 33731848
2021 T-BET and EOMES Accelerate and Enhance Functional Differentiation of Human Natural Killer Cells. Frontiers in immunology 43 34630413
2019 GATA6 Cooperates with EOMES/SMAD2/3 to Deploy the Gene Regulatory Network Governing Human Definitive Endoderm and Pancreas Formation. Stem cell reports 43 30629940
2021 EOMES and IL-10 regulate antitumor activity of T regulatory type 1 CD4+ T cells in chronic lymphocytic leukemia. Leukemia 39 33526861
2012 Tbr2 deficiency in mitral and tufted cells disrupts excitatory-inhibitory balance of neural circuitry in the mouse olfactory bulb. The Journal of neuroscience : the official journal of the Society for Neuroscience 39 22745484
2019 Eomes transcription factor is required for the development and differentiation of invariant NKT cells. Communications biology 38 31044175
2016 Predominance of weakly cytotoxic, T-betLowEomesNeg CD8+ T-cells in human gastrointestinal mucosa: implications for HIV infection. Mucosal immunology 36 27827375
2013 Pax6 regulates Tbr1 and Tbr2 expressions in olfactory bulb mitral cells. Molecular and cellular neurosciences 36 23353076
2017 The Tbr2 Molecular Network Controls Cortical Neuronal Differentiation Through Complementary Genetic and Epigenetic Pathways. Cerebral cortex (New York, N.Y. : 1991) 35 27600842
2017 Eomes expression reports the progressive differentiation of IFN-γ-producing Th1-like γδ T cells. European journal of immunology 35 28386934
2016 Effect of CMV and Aging on the Differential Expression of CD300a, CD161, T-bet, and Eomes on NK Cell Subsets. Frontiers in immunology 34 27872625
2015 Evolutionary origin of Tbr2-expressing precursor cells and the subventricular zone in the developing cortex. The Journal of comparative neurology 34 26267763
2020 Eomes broadens the scope of CD8 T-cell memory by inhibiting apoptosis in cells of low affinity. PLoS biology 32 32182234
2019 The role of Eomes in human CD4 T cell differentiation: A question of context. European journal of immunology 32 30536524
2018 The Epigenetic Factor Landscape of Developing Neocortex Is Regulated by Transcription Factors Pax6→ Tbr2→ Tbr1. Frontiers in neuroscience 32 30186101
2014 Checkpoint blockade immunotherapy relies on T-bet but not Eomes to induce effector function in tumor-infiltrating CD8+ T cells. Cancer immunology research 32 25516478
2018 EOMES-positive CD4+ T cells are increased in PTPN22 (1858T) risk allele carriers. European journal of immunology 31 29388193
2021 Antigen-driven PD-1+ TOX+ BHLHE40+ and PD-1+ TOX+ EOMES+ T lymphocytes regulate juvenile idiopathic arthritis in situ. European journal of immunology 30 33296081
2018 Cardiogenic programming of human pluripotent stem cells by dose-controlled activation of EOMES. Nature communications 30 29382828
2020 Eomes promotes esophageal carcinoma progression by recruiting Treg cells through the CCL20-CCR6 pathway. Cancer science 28 33113266
2019 Liver-Derived TGF-β Maintains the EomeshiTbetlo Phenotype of Liver Resident Natural Killer Cells. Frontiers in immunology 28 31333651
2017 Functional characterisation of cis-regulatory elements governing dynamic Eomes expression in the early mouse embryo. Development (Cambridge, England) 28 28174238
2013 Generation and characterization of a tamoxifen-inducible Eomes(CreER) mouse line. Genesis (New York, N.Y. : 2000) 28 23897762
2019 Extrapituitary prolactin promotes generation of Eomes-positive helper T cells mediating neuroinflammation. Proceedings of the National Academy of Sciences of the United States of America 27 31570595
2014 CB1 Cannabinoid Receptor-Dependent Activation of mTORC1/Pax6 Signaling Drives Tbr2 Expression and Basal Progenitor Expansion in the Developing Mouse Cortex. Cerebral cortex (New York, N.Y. : 1991) 27 24610119
2014 T-bet:Eomes balance, effector function, and proliferation of cytomegalovirus-specific CD8+ T cells during primary infection differentiates the capacity for durable immune control. Journal of immunology (Baltimore, Md. : 1950) 27 25339676
2023 Eomes restricts Brachyury functions at the onset of mouse gastrulation. Developmental cell 26 37633271
2020 Eomes Expression Defines Group 1 Innate Lymphoid Cells During Metastasis in Human and Mouse. Frontiers in immunology 24 32625207
2013 Roles of CDX2 and EOMES in human induced trophoblast progenitor cells. Biochemical and biophysical research communications 24 23313847
2018 Eomes partners with PU.1 and MITF to Regulate Transcription Factors Critical for osteoclast differentiation. iScience 23 30634169
2022 Multiple Sclerosis CSF Is Enriched With Follicular T Cells Displaying a Th1/Eomes Signature. Neurology(R) neuroimmunology & neuroinflammation 22 36266053
2021 Eomes Impedes Durable Response to Tumor Immunotherapy by Inhibiting Stemness, Tissue Residency, and Promoting the Dysfunctional State of Intratumoral CD8+ T Cells. Frontiers in cell and developmental biology 22 33553191
2021 Characterization of Tbr2-expressing retinal ganglion cells. The Journal of comparative neurology 22 34245014
2016 Reciprocal regulation of BMF and BIRC5 (Survivin) linked to Eomes overexpression in colorectal cancer. Cancer letters 21 27539959
2013 Down-regulation of interferon tau gene transcription with a transcription factor, EOMES. Molecular reproduction and development 21 23606646
2023 Eomesodermin-expressing type 1 regulatory (EOMES+ Tr1)-like T cells: Basic biology and role in immune-mediated diseases. European journal of immunology 19 36653901
2024 Eomes expression identifies the early bone marrow precursor to classical NK cells. Nature immunology 18 38871999
2023 Eomes is sufficient to regulate IL-10 expression and cytotoxic effector molecules in murine CD4+ T cells. Frontiers in immunology 18 36756120
2020 Down-regulation of EOMES drives T-cell exhaustion via abolishing EOMES-mediated repression of inhibitory receptors of T cells in liver cancer. Journal of cellular and molecular medicine 18 33325636
2020 Perinatal IL-1β-induced inflammation suppresses Tbr2+ intermediate progenitor cell proliferation in the developing hippocampus accompanied by long-term behavioral deficits. Brain, behavior, & immunity - health 18 34589867

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