Affinage

CACNA2D1

Voltage-dependent calcium channel subunit alpha-2/delta-1 · UniProt P54289

Length
1103 aa
Mass
124.6 kDa
Annotated
2026-06-09
100 papers in source corpus 52 papers cited in narrative 52 extracted findings
Cross-family judge vs UniProt: Affinage preferred faithfulness: 6/6 claims corpus-supported (100%)

Mechanistic narrative

Synthesis pass · prose summary of the discoveries below

CACNA2D1 encodes α2δ-1, a multifunctional auxiliary subunit of voltage-gated Ca2+ channels that also operates as a synaptic organizer and glutamate-receptor regulator, with central roles in synaptogenesis, central sensitization, and cardiovascular tone (PMID:19818485, PMID:29490268, PMID:24133248). As a canonical channel subunit, α2δ-1 promotes plasma-membrane trafficking and tunes the kinetics and voltage dependence of CaV1, CaV2, and CaV3 channels: it is essential for surface CaV1.2 in vascular and pancreatic cells and for CaV2 trafficking in neurons, while shaping slow L-type current kinetics in skeletal and cardiac muscle without altering targeting in those tissues (PMID:15536090, PMID:19797702, PMID:17563358, PMID:10534405, PMID:31805307); its N-terminal R-domain is necessary and sufficient for these effects on CaV2.2 (PMID:25966687). Beyond channel chaperoning, α2δ-1 is the neuronal receptor for astrocyte-secreted thrombospondins—its VWF-A domain binds thrombospondin EGF repeats and drives postsynaptic excitatory synaptogenesis and spinogenesis via Rac1—and gabapentin antagonizes this interaction (PMID:19818485, PMID:30054448). α2δ-1 forms heteromeric complexes with NMDA and AMPA receptors through its intrinsically disordered C-terminus: it promotes synaptic NMDAR trafficking and activity in a PKC-phosphorylation-dependent manner (requiring GluN2A-S929/GluN2B-S1413), disrupts GluA1/GluA2 heteromer assembly to favor Ca2+-permeable GluA1 homotetramers, and promotes ubiquitin-proteasomal degradation of GluA3 at K861 (PMID:29490268, PMID:34252035, PMID:34289359, PMID:41129242). Under neuropathic, chemotherapy, opioid, ischemic, and viral insults, injury-induced α2δ-1 upregulation in DRG neurons drives anterograde trafficking of α2δ-1–NMDAR/AMPAR complexes to spinal synapses, amplifying nociceptive transmission and central sensitization; this upregulation is governed by reduced HDAC2 occupancy at the Cacna2d1 promoter, controlled by CK2-mediated HDAC2 phosphorylation (PMID:15456823, PMID:30431158, PMID:36257688, PMID:30839350, PMID:39357831, PMID:41129242). These same complexes underlie hypertension through augmented NMDAR and CP-AMPAR activity in hypothalamic presympathetic neurons, and the α2δ-1–dependent mechanism is the molecular target exploited by gabapentinoids (gabapentin, pregabalin), which act via Arg217 (PMID:24889613, PMID:21464332, PMID:34193557, PMID:37605933, PMID:29971791). Biallelic loss-of-function CACNA2D1 variants cause developmental epileptic encephalopathy, with the G209D missense variant abolishing surface trafficking and CaV2 channel function (PMID:35293990).

Mechanistic history

Synthesis pass · year-by-year structured walk · 15 steps
  1. 1999 Medium

    Established α2δ-1 as a functional auxiliary subunit that dramatically amplifies voltage-gated Ca2+ channel currents, defining its canonical role.

    Evidence Heterologous co-expression of full-length CACNA2 with α1A/β4 in HEK293 cells with patch-clamp

    PMID:10534405

    Open questions at the time
    • Did not resolve which domains mediate enhancement
    • Single heterologous system; no native tissue validation
  2. 2004 High

    Distinguished tissue-specific channel functions of α2δ-1, showing it sets slow L-type current kinetics in muscle without governing channel targeting, and that injured-DRG-derived spinal α2δ-1 upregulation causally drives central sensitization.

    Evidence siRNA in dysgenic myotubes with patch clamp; dorsal rhizotomy plus intrathecal antisense in spinal nerve ligation model

    PMID:15456823 PMID:15536090

    Open questions at the time
    • Did not identify the trafficking partners mediating spinal upregulation
    • Downstream synaptic effectors of α2δ-1 not yet defined
  3. 2007 High

    Extended the kinetic-modulator role to cardiac muscle, where α2δ-1 loss shifts activation voltage and slows kinetics with predicted impact on action potential and Ca2+ handling.

    Evidence siRNA in reconstituted dysgenic muscle cells with patch clamp and computational ventricular myocyte modeling

    PMID:17563358

    Open questions at the time
    • Physiological consequences modeled, not measured in intact heart
  4. 2009 High

    Reframed α2δ-1 beyond a channel subunit by identifying it as the thrombospondin receptor driving excitatory CNS synaptogenesis, and showed pregabalin's antiallodynic action works by impairing α2δ-1 anterograde trafficking.

    Evidence Domain-mapping pulldown/Co-IP and synaptogenesis assays; EM/light-microscopy localization with chronic pregabalin in nerve ligation

    PMID:19339603 PMID:19818485

    Open questions at the time
    • Downstream signaling from thrombospondin–α2δ-1 binding not yet defined
    • Mechanism linking trafficking block to allodynia incomplete
  5. 2009 High

    Demonstrated α2δ-1 is essential for CaV1.2 surface expression and myogenic tone in vascular myocytes, linking it to cardiovascular function.

    Evidence shRNA knockdown with surface biotinylation, patch clamp, and pressurized artery assays

    PMID:19797702

    Open questions at the time
    • Mechanism of α2δ-1-dependent CaV1.2 surface delivery not resolved
  6. 2011 High

    Established that α2δ-1 ligand binding (via Arg217) functionally mediates gabapentinoid suppression of glutamate release and anxiolysis, and probed how membrane anchoring and lipid-raft targeting relate to current enhancement.

    Evidence α2δ-1 R217A and R279A point-mutant mice with enantioselective pharmacology and behavior; GPI/TM chimera fractionation

    PMID:21464332 PMID:21558437 PMID:21695204

    Open questions at the time
    • Raft-targeting upstream sequences not mapped
    • Link between glutamate-release modulation and channel trafficking incomplete
  7. 2012 Medium

    Questioned the requirement for membrane anchoring in current enhancement by showing C-terminally truncated, largely secreted α2δ-1 still augments CaV2.1 currents.

    Evidence Truncation mutagenesis with heterologous patch clamp and detergent-resistant membrane fractionation

    PMID:22869375

    Open questions at the time
    • Single-lab study; mechanism of anchor-independent enhancement unresolved
  8. 2013 High

    Genetic ablation confirmed α2δ-1 is required for CaV2.2 abundance and nociceptive sensitivity and is necessary for pregabalin's analgesic action; a nerve-injury-induced splice variant was found to retain channel function but lose gabapentin affinity.

    Evidence α2δ-1 KO mice with synaptosome immunoblotting, DRG patch clamp, behavior; RT-PCR splice variant with radioligand binding

    PMID:24133248 PMID:24315988

    Open questions at the time
    • Functional significance of gabapentin-resistant splice variant in vivo not established
  9. 2014 High

    Characterized the surface architecture of the α2δ-1–CaV2.2 association, identified USP2-45-dependent deubiquitylation of the channel complex, and linked α2δ-1 to vascular and addiction contexts.

    Evidence Antigen-retrieval surface labeling with gabapentin-insensitive mutant; Co-IP/biotinylation with USP2-45; SHR vascular assays; cocaine self-administration with NAc electrophysiology

    PMID:22949532 PMID:24889613 PMID:24948814 PMID:25366495

    Open questions at the time
    • USP2-45 finding single-lab; physiological role uncertain
    • Addiction and hypertension findings Medium-confidence, single-lab
  10. 2018 High

    Identified the α2δ-1–NMDAR interaction as a core mechanism: α2δ-1 binds NMDARs via its C-terminus, drives their synaptic trafficking and hyperactivity across spinal pain, ischemia, hypertension, striatal LTP, and chemotherapy contexts, conserved in human tissue and disruptable by a C-terminus peptide.

    Evidence Reciprocal Co-IP in rodent and human tissue, Cacna2d1 KO, electrophysiology, C-terminus interfering peptide across multiple disease models

    PMID:29490268 PMID:29921713 PMID:29971791 PMID:30054448 PMID:30104341 PMID:30355118 PMID:30355732 PMID:30431158

    Open questions at the time
    • Structural basis of the C-terminal NMDAR interaction not resolved
    • How α2δ-1 selects NMDAR cargo for synaptic delivery unclear
  11. 2021 High

    Defined the regulatory and AMPAR arms: PKC phosphorylation (GluN2A-S929/GluN2B-S1413) gates α2δ-1-driven NMDAR trafficking, and α2δ-1 disrupts GluA1/GluA2 assembly to promote Ca2+-permeable AMPARs, extending the mechanism to multiple neuropathies.

    Evidence Phosphoproteomics with site-directed mutagenesis, Co-IP, ER fractionation, electrophysiology, KO mice across RTX and nerve injury

    PMID:34193557 PMID:34252035 PMID:34252037 PMID:34289359

    Open questions at the time
    • Kinase/phosphatase balance controlling the switch incompletely mapped
    • Whether NMDAR and AMPAR complexes are spatially distinct unclear
  12. 2022 High

    Linked CACNA2D1 to human disease and established epigenetic control of its expression: biallelic loss-of-function causes developmental epileptic encephalopathy, while HDAC2 occupancy at the Cacna2d1 promoter constitutively represses transcription and is lost after nerve injury.

    Evidence Patient cells with G209D reconstitution and trafficking assays; ChIP with conditional Hdac2 KO and double-KO epistasis; opioid models with KO mice

    PMID:35038178 PMID:35293990 PMID:36222452 PMID:36257688 PMID:36379705

    Open questions at the time
    • Full spectrum of disease-causing variants not catalogued
    • Upstream signals triggering HDAC2 displacement only partly defined
  13. 2024 High

    Identified CK2-mediated HDAC2 phosphorylation (S394) as the upstream switch derepressing Cacna2d1 after injury, and showed calcineurin and CK2 antagonistically control α2δ-1-dependent CP-AMPAR phenotypes in spinal excitatory neurons.

    Evidence Co-IP, ChIP-qPCR, CK2 inhibitor CX-4945, phosphosite immunoblotting, electrophysiology in genetically identified neurons with KO

    PMID:38886057 PMID:39357831

    Open questions at the time
    • What activates CK2 after nerve injury is unresolved
  14. 2025 High

    Resolved the molecular basis of α2δ-1-driven CP-AMPAR formation, showing α2δ-1 specifically promotes ubiquitin-proteasomal degradation of GluA3 at K861, and placed the mechanism downstream of tonic inhibitory and mGluR5 control.

    Evidence Ubiquitination assays with K861 mutagenesis, intrathecal Gria3 gene delivery, KO mice, electrophysiology with disinhibition and mGluR5 blockade

    PMID:41006062 PMID:41129242

    Open questions at the time
    • Identity of the E3 ligase acting on GluA3 not established
    • How α2δ-1 couples to the proteasomal machinery unclear
  15. 2026 High

    Extended the α2δ-1–NMDAR mechanism to HIV-associated neuropathic pain and identified an lncRNA/hnRNPA2B1 axis stabilizing Cacna2d1 mRNA, broadening the regulatory landscape of α2δ-1 expression.

    Evidence Co-IP, KO and conditional Grin1 KO with electrophysiology and behavior (gp120); RNA pull-down and RIP with lncRNA knockdown/overexpression

    PMID:41864512 PMID:42225412

    Open questions at the time
    • lncRNA finding Medium-confidence, single-lab
    • Generalizability of mRNA-stabilization axis across tissues unknown

Open questions

Synthesis pass · forward-looking unresolved questions
  • How α2δ-1 mechanistically discriminates among its many partners (channels, thrombospondins, NMDARs, AMPARs) at distinct subcellular sites, and the structural basis of the C-terminal receptor interactions, remain unresolved.
  • No experimental structure of α2δ-1 bound to NMDARs or AMPARs
  • Mechanism partitioning channel-dependent versus channel-independent functions undefined
  • E3 ligase and full trafficking machinery not identified

Mechanism profile

Synthesis pass · controlled-vocabulary classification · explore literature graph →
Molecular activity
GO:0098772 molecular function regulator activity 4 GO:0060090 molecular adaptor activity 3 GO:0001618 virus receptor activity 1 GO:0008289 lipid binding 1
Localization
GO:0005783 endoplasmic reticulum 4 GO:0005886 plasma membrane 4 GO:0005576 extracellular region 1
Pathway
R-HSA-112316 Neuronal System 4 R-HSA-1643685 Disease 4 R-HSA-1266738 Developmental Biology 2 R-HSA-162582 Signal Transduction 2
Complex memberships
voltage-gated Ca2+ channel complex (CaV2.2/β/α2δ-1)α2δ-1–AMPAR complexα2δ-1–NMDAR complex

Evidence

Reading pass · 52 per-paper findings extracted from the source corpus
Year Finding Method Journal Conf PMIDs
2009 α2δ-1 is the neuronal receptor for astrocyte-secreted thrombospondin that promotes excitatory CNS synaptogenesis. The VWF-A domain of α2δ-1 interacts with the EGF-like repeats common to all thrombospondins. α2δ-1 overexpression increases synaptogenesis in vitro and in vivo, and postsynaptic α2δ-1 is required for thrombospondin- and astrocyte-induced synapse formation. Gabapentin antagonizes thrombospondin binding to α2δ-1 and inhibits excitatory synapse formation. Co-immunoprecipitation, domain-mapping pulldown, overexpression/knockdown in vitro and in vivo synaptogenesis assays, gabapentin competition binding Cell High 19818485
2009 In a neuropathic pain model, α2δ-1 is increased in the endoplasmic reticulum of DRG somata and in presynaptic terminals in the dorsal horn. Pregabalin's antiallodynic effect is associated with impaired anterograde trafficking of α2δ-1, reducing its plasma membrane expression in presynaptic terminals without affecting endocytosis or DRG mRNA/protein levels. Light- and electron-microscopy immunolocalization, spinal nerve ligation rat model, chronic pregabalin treatment, in vitro plasma membrane expression assay with endocytosis measurement The Journal of neuroscience High 19339603
2018 α2δ-1 forms a heteromeric complex with NMDA receptors in rodent and human spinal cords. The interaction occurs predominantly through the C terminus of α2δ-1 and promotes surface trafficking and synaptic targeting of NMDARs. Cacna2d1 overexpression potentiates presynaptic and postsynaptic NMDAR activity; knockdown or ablation normalizes nerve-injury-increased synaptic NMDAR activity. Gabapentin or an α2δ-1 C-terminus interfering peptide normalizes NMDAR synaptic targeting. Co-immunoprecipitation, Cacna2d1 overexpression/knockdown/knockout, electrophysiological recordings, C-terminus peptide disruption assay, subcellular fractionation Cell reports High 29490268
2004 Spinal dorsal horn α2δ-1 upregulation after nerve injury derives from injured DRG neurons (presynaptic origin), demonstrated by dorsal rhizotomy abolishing spinal α2δ-1 upregulation. Intrathecal α2δ-1 antisense oligonucleotides blocked dorsal horn upregulation and tactile allodynia, establishing a causal role in central sensitization. Dorsal rhizotomy, intrathecal antisense oligonucleotides, spinal nerve ligation model, immunohistochemistry, behavioral allodynia testing The Journal of neuroscience High 15456823
2018 Postsynaptic (but not presynaptic) α2δ-1 is required and sufficient for TSP-induced synaptogenesis in vitro and spine formation in vivo in the mouse cortex. Global or cell-specific loss of α2δ-1 causes profound deficits in excitatory synapse numbers, ultrastructure, and activity, and severely stunts spinogenesis. TSP-α2δ-1 interactions control synaptogenesis postsynaptically via Rac1. Conditional and global knockout mice, cell-specific rescue experiments, electron microscopy, electrophysiology, Rac1 activity assays, in vitro synaptogenesis assay The Journal of cell biology High 30054448
2014 At the cell surface, α2δ-1 is in tight physical association with the CaV2.2 α1 subunit; the association is not disrupted by gabapentin, but gabapentin reduces CaV2.2 surface expression specifically when wild-type (not gabapentin-insensitive R217A mutant) α2δ-1 is present. The antigenic epitope of α2δ-1 is occluded by CaV2.2 at the plasma membrane, revealed by antigen retrieval. Functional exofacial-tagged CaV2.2, surface antibody labeling with antigen retrieval, co-immunoprecipitation, electrophysiology, gabapentin-insensitive α2δ-1 mutant Proceedings of the National Academy of Sciences of the United States of America High 24889613
2013 α2δ-1 knockout mice show reduced CaV2.2 levels in brain and spinal cord synaptosomes, lower calcium channel current density in DRG neurons, reduced mechanical and cold sensitivity, and delayed mechanical hypersensitivity after partial sciatic nerve ligation. The ability of pregabalin to alleviate mechanical hypersensitivity is lost in α2δ-1 knockout mice. α2δ-1 knockout mice, synaptosome immunoblotting, whole-cell patch-clamp of DRG neurons, behavioral nociception testing, in vivo electrophysiology of dorsal horn neurons The Journal of neuroscience High 24133248
2004 In skeletal muscle, α2δ-1 depletion does not affect triad targeting or functional expression of CaV1 channels (α1S targeting, L-type current amplitude, voltage dependence, EC coupling Ca2+ transients) but significantly accelerates Ca2+ current kinetics (converts slowly activating to fast activating channels), identifying α2δ-1 as a major determinant of slow L-type Ca2+ current kinetics. siRNA knockdown of α2δ-1 in dysgenic myotubes and BC3H1 cells, whole-cell patch clamp, immunofluorescence, Ca2+ transient measurements, RT-PCR The Journal of biological chemistry High 15536090
2009 α2δ-1 is the only α2δ isoform in cerebral artery myocytes and is essential for plasma membrane expression of CaV1.2 α1 subunits; α2δ-1 knockdown reduces plasma-membrane CaV1.2, increases cytosolic CaV1.2, decreases intracellular Ca2+, inhibits myogenic tone, and attenuates vasodilation. Pregabalin directly inhibits CaV1.2 currents and reduces plasma membrane expression of both α2δ-1 and CaV1.2 α1. shRNA knockdown, surface biotinylation, whole-cell patch clamp, pressurized artery myogenic tone assay, pregabalin pharmacology Circulation research High 19797702
2007 In cardiac muscle, α2δ-1 depletion does not affect membrane targeting or expression of CaV1.2 but shifts voltage dependence of Ca2+ current activation by +9 mV and slows activation/inactivation kinetics ~2-fold, predicted by computer modeling to prolong action potential by 60% and double myoplasmic Ca2+ per contraction. siRNA knockdown of α2δ-1 in reconstituted dysgenic muscle cells, whole-cell patch clamp, computer modeling of cardiac ventricular myocyte electrophysiology Proceedings of the National Academy of Sciences of the United States of America High 17563358
2011 Pregabalin inhibits K+-evoked glutamate release from rat neocortical slices via the α2δ-1 subunit, demonstrated by loss of pregabalin effect in transgenic mice carrying the gabapentin-insensitive α2δ-1 R217A point mutation. The R(-)-enantiomer of pregabalin (which lacks α2δ binding) does not inhibit glutamate release. Enzyme-based microelectrode amperometry, wild-type vs. α2δ-1 R217A point mutant mice, enantioselective pharmacology, L-isoleucine competition The Journal of pharmacology and experimental therapeutics High 21464332
2011 The anxiolytic-like effects of pregabalin are mediated by binding to the α2δ-1 subunit specifically: pregabalin is inactive in α2δ-1 R217A point-mutant mice up to 320 mg/kg but retains activity in α2δ-2 R279A mutant mice. Vogel conflict test, α2δ-1 R217A and α2δ-2 R279A point-mutant knock-in mice, dose-response pharmacology The Journal of pharmacology and experimental therapeutics High 21558437
2012 α2δ-1 can enhance Ca2+ channel currents even when its membrane anchor (GPI anchor or C-terminal hydrophobic domain) is removed; the truncated α2δ-1 (α2δ-1ΔC-term) is largely secreted but some remains extrinsically associated with lipid raft fractions, and co-expression with CaV2.1/β1b still enhances calcium currents (though less than WT), questioning the requirement for membrane anchoring for current enhancement. Truncation mutagenesis, heterologous expression, whole-cell patch clamp, detergent-resistant membrane fractionation, secretion assay The Journal of biological chemistry Medium 22869375
2011 α2δ-1 targeting to lipid rafts is independent of its GPI-anchoring motif; substituting the GPI anchor with a type I TM domain (PIN-G TM) does not abolish raft localization, indicating that raft targeting is governed by upstream sequences promoting protein-protein rather than lipid-lipid interactions. GPI/TM chimera expression, density-gradient lipid raft fractionation, electrophysiology, PI-PLC cleavage assay, immunofluorescence PloS one Medium 21695204
2014 USP2-45 de-ubiquitylates both CaV1.2 and α2δ-1 subunits, reduces CaV1.2 surface expression and current amplitude, and requires α2δ-1 co-expression for its effect. α2δ-1 (but not CaV1.2 or β2) co-precipitates with USP2-45, suggesting USP2-45 binds to α2δ-1 to promote de-ubiquitylation of the channel complex. Co-immunoprecipitation, surface biotinylation, whole-cell patch clamp, co-expression in HEK293/tsA-201 cells Pflugers Archiv : European journal of physiology Medium 25366495
2015 Upregulation of α2δ-1 in DRG sensory neurons prolongs Ca2+ responses evoked by membrane depolarization via CaV2.2-mediated Ca2+ influx; the prolonged Ca2+ transients are not dependent on extracellular Ca2+ after initiation and are buffered by mitochondria preferentially activated by Ca2+ influx through CaV2.2 channels. Overexpression of α2δ-1 in rat DRG neurons, Ca2+ imaging, ω-conotoxin GVIA block of CaV2.2, mitochondrial Ca2+ uptake inhibition, removal of extracellular Ca2+ The Journal of neuroscience Medium 25878262
2013 A naturally occurring α2δ-1 splice variant (ΔA+BΔC), differentially upregulated in small non-myelinated DRG neurons after spinal nerve ligation, supports CaV2 calcium currents with unaltered biophysical properties compared to the main variant but shows significantly reduced affinity for gabapentin. RT-PCR splice variant identification, density-gradient DRG neuron separation, heterologous expression, whole-cell patch clamp, [3H]-gabapentin binding Pain Medium 24315988
2018 α-Neurexins (αNrxns) act as positive regulators of Ca2+ influx through CaV2.1 channels specifically when the α2δ-1 (not α2δ-3) auxiliary subunit is present. αNrxn triple knockout reduces presynaptic Ca2+ influx and CaV2.1 synaptic abundance, correlating with elevated surface mobility of α2δ-1 on axons. Nrxn1α co-expression with α2δ-1 (but not α2δ-3) facilitates CaV2.1 Ca2+ currents without altering kinetics. αNrxn triple knockout neurons, Ca2+ imaging, whole-cell patch clamp of recombinant CaV2.1 channels, surface mobility tracking, rescue overexpression The Journal of neuroscience High 30104341
2018 Increased α2δ-1 expression in chemotherapy (paclitaxel)-treated rats potentiates the α2δ-1-NMDAR interaction and synaptic trafficking in the spinal cord; α2δ-1 is required for paclitaxel-induced tonic activation of presynaptic NMDARs, since inhibiting α2δ-1, disrupting α2δ-1-NMDAR interaction, or α2δ-1 genetic ablation fully reverses paclitaxel-induced presynaptic NMDAR-mediated glutamate release. Co-immunoprecipitation, α2δ-1 KO mice, pregabalin treatment, C-terminus interfering peptide, spinal cord slice electrophysiology Journal of neurochemistry High 30431158
2018 Focal cerebral ischemia rapidly enhances the α2δ-1-NMDAR physical interaction in mouse brain tissue and increases α2δ-1 protein glycosylation. α2δ-1 is essential for ischemia-induced NMDAR hyperactivity; gabapentin, C-terminus-interfering peptide, or Cacna2d1 knockout abolish oxygen-glucose deprivation-induced NMDAR hyperactivity without affecting basal NMDAR currents. Co-immunoprecipitation, MCAO stroke model, hippocampal slice electrophysiology (OGD model), gabapentin pharmacology, Cacna2d1 KO mice, C-terminus peptide Stroke High 30355118
2018 The α2δ-1-NMDAR interaction in the striatum is physically conserved in rodents and humans. TBS-induced corticostriatal LTP is abolished by gabapentin, by the α2δ-1 C-terminus interfering peptide, or by Cacna2d1 knockout. TBS switches NMDARs from α2δ-1-free to α2δ-1-bound at synapses, and this is required for presynaptic and postsynaptic NMDAR activity potentiation underlying LTP. Disrupting this interaction impairs T-maze alternation and rotarod performance. Co-immunoprecipitation (rodent and human), Cacna2d1 KO mice, TBS-induced LTP electrophysiology, gabapentin and C-terminus peptide, behavioral assays The Journal of biological chemistry High 30355732
2018 α2δ-1 interacts with NMDARs in the hypothalamus of rats and humans (Co-IP). Angiotensin II increases synaptic α2δ-1-NMDAR complexes and augments presynaptic and postsynaptic NMDAR activity via AT1 receptors; these effects are abolished by pregabalin, by the α2δ-1 C-terminus peptide, or in Cacna2d1 knockout mice. Disrupting α2δ-1-NMDAR interaction blocks the sympathoexcitatory response to PVN Ang II microinjection. Co-immunoprecipitation (rodent and human hypothalamus), Cacna2d1 KO, whole-cell patch clamp of PVN neurons, pregabalin/C-terminus peptide, in vivo sympathetic nerve recording The Journal of neuroscience High 29921713
2021 PKC activation increases α2δ-1-GluN1 interaction and promotes co-trafficking of α2δ-1 and GluN1 to the cell surface. α2δ-1 is indispensable for PKC-induced surface and synaptic trafficking of NMDARs; PKC inhibition abolishes α2δ-1-coexpression-induced NMDAR potentiation. PKC-induced NMDAR potentiation requires phosphorylation of S929 on GluN2A and S1413 on GluN2B. In vivo, PMA increases NMDAR activity and synaptic α2δ-1-GluN1 complexes in wild-type but not α2δ-1 KO mice. HEK293 transfection, Co-IP, patch-clamp electrophysiology, PMA/Gö6983 pharmacology, quantitative phosphoproteomics, site-directed mutagenesis, α2δ-1 KO mice, spinal cord synaptosome fractionation The Journal of neuroscience High 34252035
2021 α2δ-1 physically interacts with both GluA1 and GluA2 AMPA receptor subunits via its C terminus, inhibits GluA1/GluA2 heteromeric assembly, and increases GluA2 retention in the endoplasmic reticulum. Nerve injury potentiates postsynaptic Ca2+-permeable AMPARs (CP-AMPARs) in the spinal dorsal horn via α2δ-1. Gabapentin or disrupting the α2δ-1-AMPAR complex fully restores GluA1/GluA2 heterotetrameric assembly and reduces synaptic CP-AMPARs. Co-immunoprecipitation, subcellular fractionation, electrophysiology (rectification index), CP-AMPAR pharmacology, α2δ-1 overexpression/KO, C-terminus peptide disruption Cell reports High 34289359
2022 HDAC2 constitutively suppresses Cacna2d1 transcription in DRG neurons by occupying the Cacna2d1 promoter. Nerve injury causes histone hyperacetylation and reduced HDAC2 enrichment at the Cacna2d1 promoter. Hdac2 conditional knockout in DRG neurons induces long-lasting pain hypersensitivity reversed by gabapentin, NMDAR blockers, or α2δ-1-NMDAR interaction disruption; this pain hypersensitivity is blunted in Cacna2d1 KO mice. ChIP-seq/ChIP-qPCR, conditional Hdac2 KO in DRG neurons, Cacna2d1 KO mice, behavioral pain testing, spinal cord electrophysiology, gabapentin/peptide pharmacology, immunoblotting The Journal of neuroscience High 36257688
2022 Biallelic loss-of-function variants in CACNA2D1 cause developmental epileptic encephalopathy. A missense variant (G209D) severely impairs α2δ-1 function: α2δ-1G209D shows strongly reduced trafficking to the cell surface and completely fails to increase CaV2 channel trafficking and function, demonstrating that α2δ-1 is non-interchangeable with other α2δ proteins for neuronal CaV2 trafficking. Patient fibroblast protein analysis, heterologous expression of α2δ-1G209D, cell surface trafficking assay, CaV2 channel electrophysiology, nonsense-mediated decay analysis Brain High 35293990
2015 The N-terminal 'R-domain' of α2δ-1 (upstream of the transmembrane region) is necessary and sufficient for its effects on CaV2.2 calcium currents and channel trafficking, identified by systematic chimeric and truncation mapping. Chimera and truncation mutagenesis, heterologous expression, whole-cell patch clamp, electrophysiology Current molecular pharmacology Medium 25966687
2016 α2δ-1 knockout reduces AP duration and firing frequency in DRG neurons, attributed to reduced Ca2+ entry during single AP stimulation particularly in the axon proximal segment. Reduced intracellular Ca2+ buffering mimics the α2δ-1 KO effect on AP duration and firing frequency. No consistent involvement of BK or SK channels in these events was detected. α2δ-1 KO mice, whole-cell patch clamp, Ca2+ imaging with single AP stimulation, Ca2+ chelation experiments Philosophical transactions of the Royal Society of London. Series B, Biological sciences Medium 27377724
1999 Full-length CACNA2 (α2δ-1) cDNA co-transfected with α1A and β4 into HEK293 cells reconstitutes Q-type Ca2+ currents and enhances current density 18-fold compared to α1A/β4 alone; the gene spans >150 kb with ≥40 exons including 2 alternatively spliced exons. Heterologous expression in HEK293 cells, whole-cell patch clamp, genomic library cloning Genomics Medium 10534405
2016 Thrombospondin-4 reduces [3H]-gabapentin binding affinity to α2δ-1 in a Mg2+-dependent manner requiring the VWF-A domain of α2δ-1. Partial Co-IP can be detected between thrombospondin-4 and α2δ-1 intracellularly; however, no co-immunoprecipitation or association could be detected between these two proteins at the cell surface when co-expressed. [3H]-gabapentin radioligand binding, Co-immunoprecipitation, cell-surface interaction assay, α2δ-1 VWF-A domain mutant Scientific reports Medium 27076051
2018 Chronic morphine treatment increases α2δ-1 protein levels in DRG and spinal cord, increases the physical α2δ-1-NMDAR interaction, and increases prevalence of α2δ-1-bound NMDARs at spinal synapses. α2δ-1 is required for morphine-induced tonic activation of presynaptic NMDARs; gabapentin or α2δ-1 C-terminus peptide reverses this, and α2δ-1 KO abolishes opioid-induced hyperalgesia and preserves morphine analgesia. Co-immunoprecipitation, synaptosome fractionation, spinal cord slice electrophysiology, gabapentin/C-terminus peptide pharmacology, Cacna2d1 KO mice, behavioral testing Anesthesiology High 30839350
2014 α2δ-1 in the nucleus accumbens is upregulated after cocaine self-administration and extinction; gabapentin preferentially reduces EPSC amplitude and increases paired-pulse ratio in NAc slices from cocaine-experienced rats; gabapentin microinjected into NAc core attenuates cocaine-primed (but not cue-induced) reinstatement without affecting locomotion. Cocaine self-administration/extinction model, immunoblotting, whole-cell patch clamp of NAc neurons, microinjection gabapentin, behavioral reinstatement testing The Journal of neuroscience Medium 24948814
2019 TCF7L2 regulates Cacna2d1/α2δ-1 expression in pancreatic beta-cells. α2δ-1 suppression decreases voltage-gated Ca2+ currents and high-glucose-evoked Ca2+ signaling by impairing CaV1.2 trafficking to the plasma membrane (CaV1.2 accumulates in recycling endosomes). α2δ-1 overexpression increases glucose-stimulated insulin secretion and rescues TCF7L2-knockdown-induced Ca2+ signaling deficit (but not reduced insulin secretion). siRNA knockdown/overexpression of α2δ-1 and TCF7L2, whole-cell patch clamp, Ca2+ imaging, insulin secretion assay, recycling endosome fractionation, confocal imaging Molecular and cellular endocrinology Medium 31805307
2018 α2δ-1 upregulation in cerebral artery myocytes in spontaneously hypertensive rats promotes surface trafficking of CaV1.2 channels, increasing CaV1.2 current density and reducing current inactivation, leading to enhanced vasoconstriction. Pregabalin normalizes surface α2δ-1:CaV1.2 ratio, CaV1.2 current, and vasoconstriction in hypertensive rats. Surface biotinylation, whole-cell patch clamp, pressurized artery myogenic tone assay, pregabalin pharmacology, immunoblotting in SHR vs. WKY rats Hypertension Medium 22949532
2020 Calcineurin inhibition (FK506) enhances physical interaction between α2δ-1 and NMDARs and their synaptic trafficking in the spinal cord. α2δ-1 is essential for FK506-induced tonic activation of presynaptic and postsynaptic NMDARs; gabapentin, C-terminus peptide, or Cacna2d1 KO abolishes FK506-induced NMDAR hyperactivity. Presynaptic NMDARs (conditional Grin1 KO in DRG) play a prominent role in calcineurin inhibitor-induced pain. Co-immunoprecipitation, synaptosome fractionation, Cacna2d1 KO, conditional Grin1 KO in DRG, spinal cord slice electrophysiology, gabapentin/C-terminus peptide, behavioral pain testing The Journal of neuroscience High 32269108
2022 α2δ-1 promotes synaptic expression of CP-AMPARs in the hypothalamus in hypertension by inhibiting GluA1/GluA2 heteromeric assembly. Co-IP shows α2δ-1 physically interacts with GluA1 and GluA2 in rat and human hypothalamus. Disrupting the α2δ-1-AMPAR interaction restores GluA1/GluA2 heteromers in the ER and normalizes inward rectification of AMPAR-EPSCs in PVN neurons in SHR. Co-immunoprecipitation (rodent and human), ER-enriched fractionation, whole-cell patch clamp (rectification index, IEM-1460 pharmacology), gabapentin/C-terminus peptide, SHR vs. WKY Journal of neurochemistry High 35038178
2024 CK2 phosphorylates HDAC2 at serine-394 following nerve injury, enhancing HDAC2-CK2 physical interaction and reducing HDAC2 enrichment at the Cacna2d1 promoter in DRG, thereby promoting Cacna2d1 transcription and α2δ-1 upregulation. CK2 inhibition (CX-4945) reverses pain hypersensitivity, HDAC2 phosphorylation, α2δ-1 upregulation, and restores HDAC2 occupancy and reduces histone acetylation (H3K9ac, H4K5ac) at the Cacna2d1 promoter. Co-immunoprecipitation, ChIP-qPCR, CX-4945 pharmacology, immunoblotting for phospho-HDAC2 (S394), histone acetylation assay, behavioral pain testing The Journal of biological chemistry High 39357831
2025 α2δ-1 (but not α2δ-2 or α2δ-3) promotes ubiquitin-proteasome-mediated degradation of GluA3 AMPA receptor subunit, reducing GluA2/GluA3 heteromers and increasing synaptic CP-AMPAR (GluA1 homotetramer) expression in the spinal cord. K861 on the GluA3 C-terminus is the key ubiquitination site. Intrathecal Gria3 gene delivery reverses nerve injury-induced pain hypersensitivity and synaptic CP-AMPARs. Co-expression experiments, electrophysiology, ubiquitination assay, site-directed mutagenesis (K861 ubiquitination site), intrathecal gene delivery, Cacna2d1 KO, pregabalin/C-terminus peptide/proteasome inhibition The Journal of clinical investigation High 41129242
2022 Repeated morphine exposure increases α2δ-1-GluN1 physical interaction and their synaptic trafficking in the nucleus accumbens (NAc). α2δ-1 is required for morphine-induced synaptic NMDAR hyperactivity in NAc medium spiny neurons; gabapentin, C-terminus peptide, or Cacna2d1 KO blocks morphine-elevated NMDAR activity. α2δ-1-bound NMDARs in the NAc core contribute to opioid-induced conditioned reward (conditioned place preference and locomotor sensitization). Co-immunoprecipitation, synaptosome fractionation, Cacna2d1 KO, whole-cell patch clamp, gabapentin/C-terminus peptide, conditioned place preference/locomotor sensitization behavioral assays Journal of neurochemistry High 36222452
2018 In neuropathic pain (peripheral nerve injury), α2δ-1 accumulation increases the contribution of L-type (CaV1.2) channels to neurotransmission in spinal lamina II; gabapentin reduces CaV1.2/α2δ-1 currents in HEK293 cells expressing CaV1.2/β4/α2δ-1 (but not CaV1.2/β4/α2δ-3) and reduces spontaneous EPSC frequency in CCI dorsal horn neurons. Whole-cell patch clamp (spinal lamina II neurons, HEK293 cells), CCI model, nitrendipine/gabapentin pharmacology, behavioral paw withdrawal Molecular pain Medium 29580153
2021 RTX-induced sensory neuropathy increases α2δ-1 expression across multiple DRG neuron subtypes and augments the α2δ-1-GluN1 interaction in spinal synaptosomes. α2δ-1 is required for RTX-induced potentiation of presynaptic NMDAR-mediated glutamate release from primary afferents; gabapentin, Cacna2d1 KO, or α2δ-1Tat peptide normalizes mEPSC frequency and evoked EPSC amplitude. RNAscope in situ hybridization, Co-immunoprecipitation, Cacna2d1 KO, conditional Grin1 KO in DRG, spinal cord electrophysiology, gabapentin/C-terminus Tat peptide, behavioral testing The Journal of neuroscience High 34252037
2021 CUMS-induced stress increases α2δ-1 and α2δ-1-GluN1 complexes in the PVN in borderline hypertensive rats; this drives augmented NMDAR activity in PVN presympathetic neurons and persistent hypertension. Gabapentin, memantine, AP-5, or α2δ-1Tat peptide microinjected into the PVN normalize blood pressure and renal sympathetic nerve activity in stressed borderline hypertensive rats. Co-immunoprecipitation, synaptosome fractionation, whole-cell patch clamp of PVN neurons, gabapentin/C-terminus peptide/AP-5, radiotelemetry blood pressure, renal sympathetic nerve recording The Journal of neuroscience High 34193557
2024 Calcineurin and CK2 antagonistically regulate synaptic CP-AMPAR phenotypes via α2δ-1 in spinal excitatory (VGluT2) neurons. FK506-induced calcineurin inhibition increases α2δ-1 interactions with GluA1 and GluA2 and reduces GluA1/GluA2 heteromers in the ER; CK2 inhibition reverses these effects. The α2δ-1-AMPAR interaction is required for calcineurin inhibitor-induced synaptic CP-AMPARs, as pregabalin, Cacna2d1 KO, or C-terminus peptide all reverse inward rectification of AMPAR-EPSCs specifically in VGluT2 neurons. Co-immunoprecipitation, ER fractionation, whole-cell patch clamp of genetically identified spinal neurons, CK2 inhibitor (CX-4945), Cacna2d1 KO, pregabalin/C-terminus peptide, behavioral pain testing The Journal of neuroscience High 38886057
2023 α2δ-1 is essential for calcineurin inhibitor (FK506)-induced synaptic NMDAR hyperactivity in PVN presympathetic neurons and sustained hypertension. FK506 increases α2δ-1, GluN1, and α2δ-1-GluN1 complex levels in PVN synaptosomes; these effects are blocked by gabapentin or C-terminus peptide. Cacna2d1 KO mice do not develop FK506-induced hypertension; gabapentin prevents FK506-induced hypertension in rats. Co-immunoprecipitation, synaptosome immunoblotting, whole-cell patch clamp of PVN neurons, Cacna2d1 KO, gabapentin/C-terminus peptide, radiotelemetry blood pressure, renal sympathetic nerve recording Circulation research High 37605933
2026 HIV-1 gp120 increases α2δ-1 and GluN1 expression in DRG and spinal cord, enhances α2δ-1-GluN1 interaction and synaptic trafficking. Gp120 selectively potentiates presynaptic and postsynaptic NMDAR activity in spinal VGluT2 (excitatory) but not VGAT (inhibitory) neurons. Gabapentin or α2δ-1 C-terminus peptide eliminates gp120-induced NMDAR hyperactivity. Cacna2d1 KO or conditional Grin1 KO in DRG neurons attenuates gp120-induced pain hypersensitivity. Co-immunoprecipitation, synaptosome fractionation, Cacna2d1 KO, conditional Grin1 KO in DRG, whole-cell patch clamp of genetically identified spinal neurons, gabapentin/C-terminus peptide, behavioral pain testing The Journal of neuroscience High 42225412
2014 Reduced α2δ-1 cell-surface expression in the BDNF-mutant ventromedial hypothalamus (VMH) contributes to hyperphagia; gabapentin infusion into wild-type VMH increases feeding and body weight gain; viral-mediated α2δ-1 rescue in BDNF-mutant VMH mitigates hyperphagia, obesity, and liver steatosis and normalizes glucose homeostasis. Whole-cell recordings in BDNF-mutant VMH neurons show normal calcium currents but reduced EPSC frequency, suggesting calcium channel-independent effects on feeding via α2δ-1-thrombospondin synaptogenic interactions. Viral vector rescue of α2δ-1 in VMH, gabapentin microinfusion, whole-cell patch clamp of VMH neurons, behavioral feeding assays, metabolic phenotyping The Journal of neuroscience Medium 24403154
2018 Cryo-EM-based computational modeling of the α2δ-1 binding site identifies that gabapentin, pregabalin, and related amino acidic drugs all access the same occluded binding pocket via the same entry path, with Arg217 as the key binding residue and Asp428 and Asp467 as additional anchoring points, consistent with prior mutagenesis data. Cryo-EM structure-based in silico protein-ligand sampling (exhaustive docking/MD), validated against existing mutagenesis data Journal of chemical information and modeling Low 30053380
2023 α2δ-1-NMDAR complexes in the hypothalamus of SHR are increased, and α2δ-1 is essential for potentiated presynaptic and postsynaptic NMDAR activity of PVN presympathetic neurons. Disrupting the α2δ-1-NMDAR association (C-terminus peptide into PVN) substantially reduces arterial blood pressure and renal sympathetic nerve discharges in SHRs. Co-immunoprecipitation, synaptosome fractionation, whole-cell patch clamp of retrograde-labeled PVN neurons, gabapentin/C-terminus peptide, in vivo blood pressure and sympathetic nerve recording, SHR vs. WKY The Journal of physiology High 29971791
2022 Brief μ-opioid receptor stimulation (DAMGO) selectively elicits long-term potentiation of primary afferent input in VGluT2 (excitatory) but not VGAT (inhibitory) dorsal horn neurons. This opioid-elicited LTP requires PKC, α2δ-1 (gabapentin, Cacna2d1 KO abolish it), and the α2δ-1-NMDAR interaction. Conditional Grin1 KO in DRG neurons diminishes DAMGO-elicited LTP and attenuates morphine-induced hyperalgesia and analgesic tolerance. Whole-cell patch clamp of genetically identified spinal neurons, Cacna2d1 KO, conditional Grin1 KO in DRG, PKC inhibitor, gabapentin/C-terminus peptide, behavioral pain testing The Journal of neuroscience High 36379705
2025 GABAergic and glycinergic tonic inhibition in the spinal cord tonically suppresses α2δ-1-dependent presynaptic and postsynaptic NMDAR activity at primary afferent→VGluT2 neuron synapses. Disinhibition-induced NMDAR hyperactivity and pain hypersensitivity require both Cacna2d1 and mGluR5, demonstrated by Cacna2d1 KO, C-terminus peptide, and mGluR5 blocker all preventing disinhibition-induced NMDAR synaptic hyperactivity in VGluT2 neurons. Whole-cell patch clamp of genetically identified spinal neurons, Cacna2d1 KO, conditional Grin1 KO in DRG, α2δ-1 C-terminus peptide, gabazine/strychnine disinhibition, mGluR5 blocker, behavioral pain testing The Journal of neuroscience High 41006062
2024 ΔFOSB transcription factor directly binds to the Cacna2d1 gene promoter (confirmed by ChIP-qPCR), and the ΔFOSB-CACNA2D1 axis mediates cigarette smoke-induced calcium overload in hippocampal neurons. ΔFOSB knockdown prevents CS-induced CACNA2D1 upregulation and calcium dysregulation; ΔFOSB overexpression mimics CS effects. ChIP-qPCR, ΔFOSB knockdown/overexpression, Ca2+ imaging, CS mouse model Ecotoxicology and environmental safety Medium 37167740
2026 lncRNA 4930544M13Rik-201 increases Cacna2d1 mRNA stability and protein expression by physically interacting with hnRNPA2B1 in trigeminal ganglion neurons; silencing 4930544M13Rik-201 or hnRNPA2B1 reduces CACNA2D1 expression and alleviates neuropathic pain, while overexpression of 4930544M13Rik-201 increases Cacna2d1 and causes allodynia. RNA pull-down, RNA immunoprecipitation, RNAscope FISH, RT-qPCR/Western blot, lncRNA knockdown/overexpression, Cacna2d1 inhibition, behavioral pain testing Brain research bulletin Medium 41864512

Source papers

Stage 0 corpus · 100 papers · ranked by NIH iCite citations
Year Title Journal Citations PMID
2009 Gabapentin receptor alpha2delta-1 is a neuronal thrombospondin receptor responsible for excitatory CNS synaptogenesis. Cell 736 19818485
2009 The increased trafficking of the calcium channel subunit alpha2delta-1 to presynaptic terminals in neuropathic pain is inhibited by the alpha2delta ligand pregabalin. The Journal of neuroscience : the official journal of the Society for Neuroscience 351 19339603
2018 The α2δ-1-NMDA Receptor Complex Is Critically Involved in Neuropathic Pain Development and Gabapentin Therapeutic Actions. Cell reports 231 29490268
2004 Spinal dorsal horn calcium channel alpha2delta-1 subunit upregulation contributes to peripheral nerve injury-induced tactile allodynia. The Journal of neuroscience : the official journal of the Society for Neuroscience 201 15456823
2001 Dorsal root ganglion neurons show increased expression of the calcium channel alpha2delta-1 subunit following partial sciatic nerve injury. Brain research. Molecular brain research 177 11687271
2018 Thrombospondin receptor α2δ-1 promotes synaptogenesis and spinogenesis via postsynaptic Rac1. The Journal of cell biology 125 30054448
2014 Functional exofacially tagged N-type calcium channels elucidate the interaction with auxiliary α2δ-1 subunits. Proceedings of the National Academy of Sciences of the United States of America 112 24889613
2013 α2δ-1 gene deletion affects somatosensory neuron function and delays mechanical hypersensitivity in response to peripheral nerve damage. The Journal of neuroscience : the official journal of the Society for Neuroscience 101 24133248
2004 The Ca2+ channel alpha2delta-1 subunit determines Ca2+ current kinetics in skeletal muscle but not targeting of alpha1S or excitation-contraction coupling. The Journal of biological chemistry 74 15536090
2009 Smooth muscle cell alpha2delta-1 subunits are essential for vasoregulation by CaV1.2 channels. Circulation research 69 19797702
2018 Increased α2δ-1-NMDA receptor coupling potentiates glutamatergic input to spinal dorsal horn neurons in chemotherapy-induced neuropathic pain. Journal of neurochemistry 65 30431158
2018 Focal Cerebral Ischemia and Reperfusion Induce Brain Injury Through α2δ-1-Bound NMDA Receptors. Stroke 64 30355118
2018 α-Neurexins Together with α2δ-1 Auxiliary Subunits Regulate Ca2+ Influx through Cav2.1 Channels. The Journal of neuroscience : the official journal of the Society for Neuroscience 63 30104341
2014 Genomic aberrations of the CACNA2D1 gene in three patients with epilepsy and intellectual disability. European journal of human genetics : EJHG 57 25074461
2011 Amperometric measurement of glutamate release modulation by gabapentin and pregabalin in rat neocortical slices: role of voltage-sensitive Ca2+ α2δ-1 subunit. The Journal of pharmacology and experimental therapeutics 54 21464332
2021 Protein Kinase C-Mediated Phosphorylation and α2δ-1 Interdependently Regulate NMDA Receptor Trafficking and Activity. The Journal of neuroscience : the official journal of the Society for Neuroscience 47 34252035
2012 Transcriptional upregulation of α2δ-1 elevates arterial smooth muscle cell voltage-dependent Ca2+ channel surface expression and cerebrovascular constriction in genetic hypertension. Hypertension (Dallas, Tex. : 1979) 46 22949532
2007 Computer modeling of siRNA knockdown effects indicates an essential role of the Ca2+ channel alpha2delta-1 subunit in cardiac excitation-contraction coupling. Proceedings of the National Academy of Sciences of the United States of America 46 17563358
2019 α2δ-1-Bound N-Methyl-D-aspartate Receptors Mediate Morphine-induced Hyperalgesia and Analgesic Tolerance by Potentiating Glutamatergic Input in Rodents. Anesthesiology 45 30839350
2021 α2δ-1 Upregulation in Primary Sensory Neurons Promotes NMDA Receptor-Mediated Glutamatergic Input in Resiniferatoxin-Induced Neuropathy. The Journal of neuroscience : the official journal of the Society for Neuroscience 44 34252037
2020 Calcineurin Inhibition Causes α2δ-1-Mediated Tonic Activation of Synaptic NMDA Receptors and Pain Hypersensitivity. The Journal of neuroscience : the official journal of the Society for Neuroscience 43 32269108
2018 The α2δ-1-NMDA receptor coupling is essential for corticostriatal long-term potentiation and is involved in learning and memory. The Journal of biological chemistry 43 30355732
2015 The upregulation of α2δ-1 subunit modulates activity-dependent Ca2+ signals in sensory neurons. The Journal of neuroscience : the official journal of the Society for Neuroscience 43 25878262
2021 α2δ-1 switches the phenotype of synaptic AMPA receptors by physically disrupting heteromeric subunit assembly. Cell reports 41 34289359
2018 α2δ-1 Is Essential for Sympathetic Output and NMDA Receptor Activity Potentiated by Angiotensin II in the Hypothalamus. The Journal of neuroscience : the official journal of the Society for Neuroscience 41 29921713
2011 Anxiolytic-like activity of pregabalin in the Vogel conflict test in α2δ-1 (R217A) and α2δ-2 (R279A) mouse mutants. The Journal of pharmacology and experimental therapeutics 41 21558437
2018 α2δ-1 couples to NMDA receptors in the hypothalamus to sustain sympathetic vasomotor activity in hypertension. The Journal of physiology 37 29971791
2022 HDAC2 in Primary Sensory Neurons Constitutively Restrains Chronic Pain by Repressing α2δ-1 Expression and Associated NMDA Receptor Activity. The Journal of neuroscience : the official journal of the Society for Neuroscience 36 36257688
2017 Systems genetics identifies a role for Cacna2d1 regulation in elevated intraocular pressure and glaucoma susceptibility. Nature communications 36 29176626
2013 Differential upregulation in DRG neurons of an α2δ-1 splice variant with a lower affinity for gabapentin after peripheral sensory nerve injury. Pain 36 24315988
2022 Biallelic CACNA2D1 loss-of-function variants cause early-onset developmental epileptic encephalopathy. Brain : a journal of neurology 35 35293990
2016 Thrombospondin-4 reduces binding affinity of [(3)H]-gabapentin to calcium-channel α2δ-1-subunit but does not interact with α2δ-1 on the cell-surface when co-expressed. Scientific reports 34 27076051
2012 Calcium currents are enhanced by α2δ-1 lacking its membrane anchor. The Journal of biological chemistry 33 22869375
2018 Calcium channel α2δ1 subunit (CACNA2D1) enhances radioresistance in cancer stem-like cells in non-small cell lung cancer cell lines. Cancer management and research 31 30464601
2024 Quercetin alleviates chronic unpredictable mild stress-induced depression-like behavior by inhibiting NMDAR1 with α2δ-1 in rats. CNS neuroscience & therapeutics 30 38615365
2017 Epileptiform activity and behavioral arrests in mice overexpressing the calcium channel subunit α2δ-1. Neurobiology of disease 29 28193459
2011 Targeting of voltage-gated calcium channel α2δ-1 subunit to lipid rafts is independent from a GPI-anchoring motif. PloS one 29 21695204
2021 α2δ-1-Dependent NMDA Receptor Activity in the Hypothalamus Is an Effector of Genetic-Environment Interactions That Drive Persistent Hypertension. The Journal of neuroscience : the official journal of the Society for Neuroscience 27 34193557
2014 Hypothalamic dysfunction of the thrombospondin receptor α2δ-1 underlies the overeating and obesity triggered by brain-derived neurotrophic factor deficiency. The Journal of neuroscience : the official journal of the Society for Neuroscience 24 24403154
2014 α2δ-1 signaling in nucleus accumbens is necessary for cocaine-induced relapse. The Journal of neuroscience : the official journal of the Society for Neuroscience 24 24948814
2014 Altered expression of the voltage-gated calcium channel subunit α₂δ-1: a comparison between two experimental models of epilepsy and a sensory nerve ligation model of neuropathic pain. Neuroscience 23 24641886
2022 Brief Opioid Exposure Paradoxically Augments Primary Afferent Input to Spinal Excitatory Neurons via α2δ-1-Dependent Presynaptic NMDA Receptors. The Journal of neuroscience : the official journal of the Society for Neuroscience 22 36379705
2020 MicroRNA-107 inhibits proliferation and invasion of laryngeal squamous cell carcinoma cells by targeting CACNA2D1 in vitro. Anti-cancer drugs 21 31725046
2016 Oxaliplatin administration increases expression of the voltage-dependent calcium channel α2δ-1 subunit in the rat spinal cord. Journal of pharmacological sciences 21 26883453
2023 Brain α2δ-1-Bound NMDA Receptors Drive Calcineurin Inhibitor-Induced Hypertension. Circulation research 20 37605933
2018 Peripheral nerve injury increases contribution of L-type calcium channels to synaptic transmission in spinal lamina II: Role of α2δ-1 subunits. Molecular pain 20 29580153
2014 miR‑107 promotes the erythroid differentiation of leukemia cells via the downregulation of Cacna2d1. Molecular medicine reports 20 25373460
2010 Novel SNPs of the bovine CACNA2D1 gene and their association with carcass and meat quality traits. Molecular biology reports 20 20585886
2020 Effect and mechanism of the CACNA2D1-CGRP pathway in osteoarthritis-induced ongoing pain. Biomedicine & pharmacotherapy = Biomedecine & pharmacotherapie 19 32570114
2014 Ubiquitin-specific protease USP2-45 acts as a molecular switch to promote α2δ-1-induced downregulation of Cav1.2 channels. Pflugers Archiv : European journal of physiology 19 25366495
2017 α2δ-1 Signaling Drives Cell Death, Synaptogenesis, Circuit Reorganization, and Gabapentin-Mediated Neuroprotection in a Model of Insult-Induced Cortical Malformation. eNeuro 18 29109971
2016 Effect of knockout of α2δ-1 on action potentials in mouse sensory neurons. Philosophical transactions of the Royal Society of London. Series B, Biological sciences 18 27377724
1996 Anesthesiologic problems in Williams syndrome: the CACNL2A locus is not involved. Human genetics 18 8707301
2011 Single nucleotide polymorphism of CACNA2D1 gene and its association with milk somatic cell score in cattle. Molecular biology reports 17 21225462
2022 α2δ-1 protein promotes synaptic expression of Ca2+ permeable-AMPA receptors by inhibiting GluA1/GluA2 heteromeric assembly in the hypothalamus in hypertension. Journal of neurochemistry 16 35038178
2018 Pushing the Limits of Computational Structure-Based Drug Design with a Cryo-EM Structure: The Ca2+ Channel α2δ-1 Subunit as a Test Case. Journal of chemical information and modeling 16 30053380
2014 Mapping breakpoints of a familial chromosome insertion (18,7) (q22.1; q36.2q21.11) to DPP6 and CACNA2D1 genes in an azoospermic male. Gene 16 24937803
2010 Genetic polymorphisms of the CACNA2D1 gene and their association with carcass and meat quality traits in cattle. Biochemical genetics 16 20549332
1999 Genomic structure and functional expression of a human alpha(2)/delta calcium channel subunit gene (CACNA2). Genomics 16 10534405
2025 α2δ-1-Linked NMDA and AMPA Receptors in Neuropathic Pain and Gabapentinoid Action. Journal of neurochemistry 15 40191897
2018 Candidate SNP of CACNA2D1 Gene Associated with Clinical Mastitis and Production Traits in Sahiwal (Bos taurus indicus) and Karan Fries (Bos taurus taurus × Bos taurus indicus). Animal biotechnology 15 29463160
2022 Functions and Clinical Significance of CACNA2D1 in Gastric Cancer. Annals of surgical oncology 14 35445337
2021 The α2δ-1/NMDA receptor complex is involved in brain injury after intracerebral hemorrhage in mice. Annals of clinical and translational neurology 14 34032393
2019 The TCF7L2-dependent high-voltage activated calcium channel subunit α2δ-1 controls calcium signaling in rodent pancreatic beta-cells. Molecular and cellular endocrinology 13 31805307
2022 α2δ-1 protein drives opioid-induced conditioned reward and synaptic NMDA receptor hyperactivity in the nucleus accumbens. Journal of neurochemistry 12 36222452
2015 Upregulation of α₂δ-1 Calcium Channel Subunit in the Spinal Cord Contributes to Pelvic Organ Cross-Sensitization in a Rat Model of Experimentally-Induced Endometriosis. Neurochemical research 11 25935199
2023 The α2δ-1-NMDA receptor complex and its potential as a therapeutic target for ischemic stroke. Frontiers in neurology 10 37153659
2021 Calcium Channel Subunit α2δ-1 as a Potential Biomarker Reflecting Illness Severity and Neuroinflammation in Patients with Acute Ischemic Stroke. Journal of stroke and cerebrovascular diseases : the official journal of National Stroke Association 10 34049015
2021 The α2δ-1-NMDAR1 interaction in the trigeminal ganglion contributes to orofacial ectopic pain following inferior alveolar nerve injury. Brain research bulletin 9 33811955
2014 The perfect storm? Histiocytoid cardiomyopathy and compound CACNA2D1 and RANGRF mutation in a baby. Cardiology in the young 9 24438356
1996 Influence of protein conditioning films on binding of a bacterial polysaccharide adhesin from Hyphomonas MHS-3. Biofouling 9 22115100
2024 Calcineurin and CK2 Reciprocally Regulate Synaptic AMPA Receptor Phenotypes via α2δ-1 in Spinal Excitatory Neurons. The Journal of neuroscience : the official journal of the Society for Neuroscience 8 38886057
2022 Development of a PET radioligand for α2δ-1 subunit of calcium channels for imaging neuropathic pain. European journal of medicinal chemistry 7 36031695
2021 Aberrant Axo-Axonic Synaptic Reorganization in the Phosphorylated L1-CAM/Calcium Channel Subunit α2δ-1-Containing Central Terminals of Injured c-Fibers in the Spinal Cord of a Neuropathic Pain Model. eNeuro 7 33500315
2024 Exercise training affects calcium ion transport by downregulating the CACNA2D1 protein to reduce hypertension-induced myocardial injury in mice. iScience 6 38495825
2023 Cigarette smoke triggers calcium overload in mouse hippocampal neurons via the ΔFOSB-CACNA2D1 axis to impair cognitive performance. Ecotoxicology and environmental safety 6 37167740
2024 CACNA2D1 regulates the progression and influences the microenvironment of colon cancer. Journal of gastroenterology 5 38536483
2023 Cherry leaf decoction inhibits NMDAR expression and thereby ameliorates CUMS- induced depression-like behaviors through downregulation of α2δ-1. Heliyon 5 38034773
2022 The Novel Gabapentinoid Mirogabalin Prevents Upregulation of α2δ-1 Subunit of Voltage-Gated Calcium Channels in Spinal Dorsal Horn in a Rat Model of Spinal Nerve Ligation. Drug research 5 36216339
2015 The R-Domain: Identification of an N-terminal Region of the α2δ-1 Subunit Which is Necessary and Sufficient for its Effects on Cav2.2 Calcium Currents. Current molecular pharmacology 5 25966687
2024 Nerve injury augments Cacna2d1 transcription via CK2-mediated phosphorylation of the histone deacetylase HDAC2 in dorsal root ganglia. The Journal of biological chemistry 4 39357831
2020 Sex-specific Effects of α2δ-1 in the Ventromedial Hypothalamus of Female Mice Controlling Glucose and Lipid Balance. Endocrinology 4 32337532
2024 The role of voltage-gated calcium channel α2δ-1 in the occurrence and development in myofascial orofacial pain. BMC oral health 3 38735923
2022 Role of the Ca2+ channel α2δ-1 auxiliary subunit in proliferation and migration of human glioblastoma cells. PloS one 3 36520928
2025 Association of the CACNA2D1 gene with milk yield and milk quality traits in Holstein cattle. The Journal of dairy research 2 40383546
2025 AI-Driven Drug Target Screening Platform Identified Oncogene CACNA2D1 Activated by Enhancer Infestation in Epstein-Barr Virus-Associated Nasopharyngeal Carcinoma. International journal of molecular sciences 2 40429844
2025 Tonic GABA and Glycine Inhibition Control Pain Hypersensitivity via Limiting α2δ-1- and mGluR5-Dependent NMDA Receptor Activity at Primary Afferent→Excitatory Neuron Synapses. The Journal of neuroscience : the official journal of the Society for Neuroscience 2 41006062
2025 Spinal α2δ-1 induces GluA3 degradation to regulate assembly of calcium-permeable AMPA receptors and pain hypersensitivity. The Journal of clinical investigation 2 41129242
2024 Towards Multitargeted Ligands as Pain Therapeutics: Dual Ligands of the Cavα2δ-1 Subunit of Voltage-Gated Calcium Channel and the μ-Opioid Receptor. ChemMedChem 2 38230842
2024 Influence of the calcium voltage-gated channel auxiliary subunit (CACNA2D1) absence on intraocular pressure in mice. Experimental eye research 1 38373629
2022 [A novel cell tool for α2δ-1-NMDAR target-based analgesic drug discovery]. Sheng wu gong cheng xue bao = Chinese journal of biotechnology 1 35355481
2020 [MicroRNA-107 inhibits the proliferation and invasion of laryngeal squamous cell carcinoma cells by targeting CACNA2D1]. Lin chuang er bi yan hou tou jing wai ke za zhi = Journal of clinical otorhinolaryngology head and neck surgery 1 33254297
2017 Dominance effects estimation of TLR4 and CACNA2D1 genes for health and production traits using logistic regression. Journal of genetics 1 29321363
2026 LncRNA 4930544M13Rik-201 regulates CACNA2D1 expression via interacting with hnRNPA2B1 to promote neuropathic pain following nerve injury. Brain research bulletin 0 41864512
2026 The Correlation Between Plasma CACNA2D1 Protein Concentration and the Severity of Coronary Heart Disease. Journal of cardiovascular development and disease 0 42188100
2026 HIV-1 gp120 induces nociceptive hypersensitivity via α2δ-1-bound NMDA receptors at primary afferent-excitatory neuron synapses. The Journal of neuroscience : the official journal of the Society for Neuroscience 0 42225412
2025 Unraveling the Role of α2δ-1 in Cerebral Hemorrhage: Calcium Overload, Endoplasmic Reticulum Stress, and Microglial Apoptosis. Brain and behavior 0 40329779
2025 α2δ-1-NMDAR1 complex in the hypothalamic paraventricular nucleus mediates anxiety-induced sympathetic hyperactivity. Clinical autonomic research : official journal of the Clinical Autonomic Research Society 0 41219589
2024 Scavenger Receptor Class B Type I Modulates Epileptic Seizures and Receptor α2δ-1 Expression. Neurochemical research 0 39017956
2024 Novel heterozygous mutation of CACNA2D1 gene in a Chinese family with arrhythmia. BMC cardiovascular disorders 0 39354346

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